"Darwin and His Influence" by Joseph Ritchie (2014)

In this passage, “selection” refers to traits that are selected for and passed on to later generations, and “species” refers to organisms that share a common ancestor and can produce viable offspring with one another.

     Early in the nineteenth century, scientists sought to understand the differences in the earth’s flora and fauna from their archeological ancestors. The prevailing view at the time was that the differences between current and previous species were unremarkable deviations from their Platonic ideal forms. This theory hinged upon the ideals of the religious-based “created kinds” theory, which state that individuals of today are products of the organisms that were present at the earth’s creation, the result of an intelligent designer. Furthermore, these individuals believed that the differences between organisms could be explained by unseen geological and astrological forces acting on organisms slowly, throughout time. Other scientists also believed that individuals had the ability to change within their lifetimes and pass on traits to their offspring efficiently and quickly through a single generation.

     Charles Darwin and other biologists, such as Alfred Wallace, were not greatly influenced by these views and hypotheses. Their propositions stated that species evolve over many generations, due to the selective pressures of their given environments. This evolution could result in the generation of divergent traits, as well as speciation and separation from the original ancestral species. The concept that organisms were not finite or present since creation was very controversial to the scientists of the period. Some saw such an idea as unsupportable, while others perceived it as heretical and fanatical.

     Darwin set out to find support for his theory through his work, On the Origins of Species by Means of Natural Selection. He was influenced by archeological discoveries of species, which appeared to have vastly different physiological appearances from present-day organisms. Darwin decided to sail around the world on a Royal Navy ship named the H.M.S. Beagle. During his travels, he was taken to the Pacific islands of the Galapagos archipelago. The volcanic islands followed a patterned distribution on either side of the Equator. The landscapes of each island varied, with different observable flora and fauna. Through scientific observations, Darwin noticed subtle variations of finches on different islands. Some finches had large hard beaks, while others had slender beaks. Beaks were differentiated from island to island. After careful study, Darwin noticed that the beaks seemed to match the food source on each island. The large beaks were specialized for breaking open hard-shelled nuts, while the slender beaks were specialized for eating certain fruits that were abundant. Darwin hypothesized that an ancestral species of finch landed on the islands, and that over generations they became adapted to the locally abundant food sources. 

     Darwin compiled multiple instances of natural selection and incorporated discoveries made by archeologists and physiologists. He surmised that species evolve over time due to the selective pressures of their respective habitats. These events occur slowly over many generations. Each species selects for advantageous traits among its members. Over time, traits selected as advantageous by environmental pressures and stressors become commonplace in the species. This niche-forming process specializes species by rewarding those with traits most suitable for reproductive success. These traits may progress into speciation of the original species, which results in the eventual development of an entirely new species. Darwin’s theory was met with opposition at the time of its publication, and the theory of evolution remains a controversial topic in several arenas of debate.

According to evidence in the third paragraph, finches that consume nuts were more likely to possess what kind of beak?

"Darwin and His Influence" by Joseph Ritchie (2014)

In this passage, “selection” refers to traits that are selected for and passed on to later generations, and “species” refers to organisms that share a common ancestor and can produce viable offspring with one another.

     Early in the nineteenth century, scientists sought to understand the differences in the earth’s flora and fauna from their archeological ancestors. The prevailing view at the time was that the differences between current and previous species were unremarkable deviations from their Platonic ideal forms. This theory hinged upon the ideals of the religious-based “created kinds” theory, which state that individuals of today are products of the organisms that were present at the earth’s creation, the result of an intelligent designer. Furthermore, these individuals believed that the differences between organisms could be explained by unseen geological and astrological forces acting on organisms slowly, throughout time. Other scientists also believed that individuals had the ability to change within their lifetimes and pass on traits to their offspring efficiently and quickly through a single generation.

     Charles Darwin and other biologists, such as Alfred Wallace, were not greatly influenced by these views and hypotheses. Their propositions stated that species evolve over many generations, due to the selective pressures of their given environments. This evolution could result in the generation of divergent traits, as well as speciation and separation from the original ancestral species. The concept that organisms were not finite or present since creation was very controversial to the scientists of the period. Some saw such an idea as unsupportable, while others perceived it as heretical and fanatical.

     Darwin set out to find support for his theory through his work, On the Origins of Species by Means of Natural Selection. He was influenced by archeological discoveries of species, which appeared to have vastly different physiological appearances from present-day organisms. Darwin decided to sail around the world on a Royal Navy ship named the H.M.S. Beagle. During his travels, he was taken to the Pacific islands of the Galapagos archipelago. The volcanic islands followed a patterned distribution on either side of the Equator. The landscapes of each island varied, with different observable flora and fauna. Through scientific observations, Darwin noticed subtle variations of finches on different islands. Some finches had large hard beaks, while others had slender beaks. Beaks were differentiated from island to island. After careful study, Darwin noticed that the beaks seemed to match the food source on each island. The large beaks were specialized for breaking open hard-shelled nuts, while the slender beaks were specialized for eating certain fruits that were abundant. Darwin hypothesized that an ancestral species of finch landed on the islands, and that over generations they became adapted to the locally abundant food sources. 

     Darwin compiled multiple instances of natural selection and incorporated discoveries made by archeologists and physiologists. He surmised that species evolve over time due to the selective pressures of their respective habitats. These events occur slowly over many generations. Each species selects for advantageous traits among its members. Over time, traits selected as advantageous by environmental pressures and stressors become commonplace in the species. This niche-forming process specializes species by rewarding those with traits most suitable for reproductive success. These traits may progress into speciation of the original species, which results in the eventual development of an entirely new species. Darwin’s theory was met with opposition at the time of its publication, and the theory of evolution remains a controversial topic in several arenas of debate.

According to the passage, why did Darwin seek passage on the H.M.S. Beagle?

Adapted from The Evolutionist at Large by Grant Allen (1881)

I am engaged in watching a brigade of ants out on foraging duty, and intent on securing for the nest three whole segments of a deceased earthworm. They look for all the world like those busy companies one sees in the Egyptian wall paintings, dragging home a huge granite colossus by sheer force of bone and sinew. Every muscle in their tiny bodies is strained to the utmost as they pry themselves laboriously against the great boulders that strew the path, and that are known to our Brobdingnagian intelligence as grains of sand. Besides the workers themselves, a whole battalion of stragglers runs to and fro upon the broad line that leads to the headquarters of the community. The province of these stragglers, who seem so busy doing nothing, probably consists in keeping communications open, and encouraging the sturdy pullers by occasional relays of fresh workmen. I often wish that I could for a while get inside those tiny brains, and see, or rather smell, the world as ants do. For there can be little doubt that to these brave little carnivores here the universe is chiefly known as a collective bundle of odors, simultaneous or consecutive. As our world is mainly a world of visible objects, theirs, I believe, is mainly a world of olfactible things.

In the head of every one of these little creatures is something that we may fairly call a brain. Of course most insects have no real brains; the nerve-substance in their heads is a mere collection of ill-arranged ganglia, directly connected with their organs of sense. Whatever man may be, an earwig at least is a conscious, or rather a semi-conscious, automaton. He has just a few knots of nerve cells in his little pate, each of which leads straight from his dim eye or his vague ear or his indefinite organs of taste; and his muscles obey the promptings of external sensations without possibility of hesitation or consideration, as mechanically as the valve of a steam engine obeys the governor balls. The poor soul's intellect is wholly deficient, and the senses alone make up all that there is of him, subjectively considered. But it is not so with the highest insects. They have something that truly answers to the real brain of men, apes, and dogs, to the cerebral hemispheres and the cerebellum that are superadded in us mammals upon the simple sense-centers of lower creatures. Besides the eye, with its optic nerve and optic perceptive organs—besides the ear, with its similar mechanism—we mammalian lords of creation have a higher and more genuine brain, that collects and compares the information given to the senses, and sends down the appropriate messages to the muscles accordingly. Now, bees and flies and ants have got much the same sort of arrangement, on a smaller scale, within their tiny heads. On top of the little knots that do duty as nerve centers for their eyes and mouths, stand two stalked bits of nervous matter, whose duty is analogous to that of our own brains. And that is why these three sorts of insects think and reason so much more intellectually than beetles or butterflies, and why the larger part of them have organized their domestic arrangements on such an excellent cooperative plan.

We know well enough what forms the main material of thought with bees and flies, and that is visible objects. For you must think about something if you think at all; and you can hardly imagine a contemplative blow-fly setting itself down to reflect, like a Hindu devotee, on the syllable Om, or on the oneness of existence. Abstract ideas are not likely to play a large part in apian consciousness. A bee has a very perfect eye, and with this eye it can see not only form, but also color, as Sir John Lubbock's experiments have shown us. The information that it gets through its eye, coupled with other ideas derived from touch, smell, and taste, no doubt makes up the main thinkable and knowable universe as it reveals itself to the apian intelligence. To ourselves and to bees alike the world is, on the whole, a colored picture, with the notions of distance and solidity thrown in by touch and muscular effort; but sight undoubtedly plays the first part in forming our total conception of things generally.

Which of the following statements about mammals is supported by the passage?

Adapted from Ice-Caves of France and Switzerland by George Forrest Browne (1865)

This account states that the cave is in the county of Thorn, among the lowest spurs of the Carpathians. The entrance, which faces the north, and is exposed to the cold winds from the snowy part of the Carpathian range, is eighteen fathoms high and nine broad; and the cave spreads out laterally, and descends to a point fifty fathoms below the entrance, where it is twenty-six fathoms in breadth, and of irregular height. Beyond this no one had at that time penetrated, on account of the unsafe footing, although many distant echoes were returned by the farther recesses of the cave; indeed, to get even so far as this, much step-cutting was necessary.

When the external frost of winter comes on, the account proceeds, the effect in the cave is the same as if fires had been lighted there: the ice melts, and swarms of flies and bats and hares take refuge in the interior from the severity of the winter. As soon as spring arrives, the warmth of winter disappears from the interior, water exudes from the roof and is converted into ice, while the more abundant supplies which pour down on to the sandy floor are speedily frozen there. In the dog-days, the frost is so intense that a small icicle becomes in one day a huge mass of ice; but a cool day promptly brings a thaw, and the cave is looked upon as a barometer, not merely feeling, but also presaging, the changes of weather. The people of the neighborhood, when employed in field-work, arrange their labour so that the mid-day meal may be taken near the cave, when they either ice the water they have brought with them, or drink the melted ice, which they consider very good for the stomach. It had been calculated that six hundred weekly carts would not be sufficient to keep the cavern free from ice. The ground above the cave is peculiarly rich in grass.

In explanation of these phenomena, Bell threw out the following suggestions, which need no comment. The earth being of itself cold and damp, the external heat of the atmosphere, by partially penetrating into the ground, drives in this native cold to the inner parts of the earth, and makes the cold there more dense. On the other hand, when the external air is cold, it draws forth towards the surface the heat there may be in the inner part of the earth, and thus makes caverns warm. In support and illustration of this view, he states that in the hotter parts of Hungary, when the people wish to cool their wine, they dig a hole two feet deep, and place in it the flagon of wine, and, after filling up the hole again, light a blazing fire upon the surface, which cools the wine as if the flagon had been laid in ice. He also suggests that possibly the cold winds from the Carpathians bring with them imperceptible particles of snow, which reach the water of the cave, and convert it into ice. Further, the rocks of the Carpathians abound in salts, nitre, alum, etc., which may, perhaps, mingle with such snowy particles, and produce the ordinary effect of the snow and salt in the artificial production of ice.

Based on the passage, the primary reason for the lack of full exploration of the cave was that __________.

Adapted from Ice-Caves of France and Switzerland by George Forrest Browne (1865)

This account states that the cave is in the county of Thorn, among the lowest spurs of the Carpathians. The entrance, which faces the north, and is exposed to the cold winds from the snowy part of the Carpathian range, is eighteen fathoms high and nine broad; and the cave spreads out laterally, and descends to a point fifty fathoms below the entrance, where it is twenty-six fathoms in breadth, and of irregular height. Beyond this no one had at that time penetrated, on account of the unsafe footing, although many distant echoes were returned by the farther recesses of the cave; indeed, to get even so far as this, much step-cutting was necessary.

When the external frost of winter comes on, the account proceeds, the effect in the cave is the same as if fires had been lighted there: the ice melts, and swarms of flies and bats and hares take refuge in the interior from the severity of the winter. As soon as spring arrives, the warmth of winter disappears from the interior, water exudes from the roof and is converted into ice, while the more abundant supplies which pour down on to the sandy floor are speedily frozen there. In the dog-days, the frost is so intense that a small icicle becomes in one day a huge mass of ice; but a cool day promptly brings a thaw, and the cave is looked upon as a barometer, not merely feeling, but also presaging, the changes of weather. The people of the neighborhood, when employed in field-work, arrange their labour so that the mid-day meal may be taken near the cave, when they either ice the water they have brought with them, or drink the melted ice, which they consider very good for the stomach. It had been calculated that six hundred weekly carts would not be sufficient to keep the cavern free from ice. The ground above the cave is peculiarly rich in grass.

In explanation of these phenomena, Bell threw out the following suggestions, which need no comment. The earth being of itself cold and damp, the external heat of the atmosphere, by partially penetrating into the ground, drives in this native cold to the inner parts of the earth, and makes the cold there more dense. On the other hand, when the external air is cold, it draws forth towards the surface the heat there may be in the inner part of the earth, and thus makes caverns warm. In support and illustration of this view, he states that in the hotter parts of Hungary, when the people wish to cool their wine, they dig a hole two feet deep, and place in it the flagon of wine, and, after filling up the hole again, light a blazing fire upon the surface, which cools the wine as if the flagon had been laid in ice. He also suggests that possibly the cold winds from the Carpathians bring with them imperceptible particles of snow, which reach the water of the cave, and convert it into ice. Further, the rocks of the Carpathians abound in salts, nitre, alum, etc., which may, perhaps, mingle with such snowy particles, and produce the ordinary effect of the snow and salt in the artificial production of ice.

The passage states that which of the following is true?

Adapted from “In Mammoth Cave” by John Burroughs (1894)

Some idea of the impression which Mammoth Cave makes upon the senses, irrespective even of sight, may be had from the fact that blind people go there to see it, and are greatly struck with it. I was assured that this is a fact. The blind seem as much impressed by it as those who have their sight. When the guide pauses at a more interesting point, or lights the scene up with a great torch or with small flares, and points out the more striking features, the blind exclaim, "How wonderful! How beautiful!" They can feel it, if they cannot see it. They get some idea of the spaciousness when words are uttered. The voice goes forth in these colossal chambers like a bird. When no word is spoken, the silence is of a kind never experienced on the surface of the earth, it is so profound and abysmal. This, and the absolute darkness, to a sighted person makes him feel as if he were face to face with the primordial nothingness. The objective universe is gone; only the subjective remains; the sense of hearing is inverted, and reports only the murmurs from within. The blind miss much, but much remains to them. The great cave is not merely a spectacle to the eye; it is a wonder to the ear, a strangeness to the smell and to the touch. The body feels the presence of unusual conditions through every pore.

Which of the following answer choices is supported by the author’s statement that inside Mammoth Cave, “The objective universe is gone”?

Adapted from Taking a Second Look: An Analysis of Genetic Markers in Species Relatedness by Joseph Ritchie (2014)

Phylogenetics is the study of genetic composition in various species and is used by evolutionary biologists to investigate similarities in the molecular sequences of proteins in varying organisms. The amino acid sequences that build proteins are used to construct mathmatical matrices that aid in determining evolutionary ties through the investigation of percentage similarities. The study of these matrices helps to expose evolutionary relationships between species that may not have the same existential characteristics.

Species adapt and evolve based on the pressures that exist in their environment. Climate, food source, and habitat availability are only a few factors that act on species adaptation. These stressors can alter the physical characteristics of organisms. This divergence in evolution has made it difficult to determine the interrelatedness of organisms by analyzing their physical characteristics alone.

For instance, looking only at physical characteristics, the ghost bat resembles a pigeon more than a spider monkey; however, phylogenetics has found that the amino acid sequences that construct the beta hemoglobin molecules of bats are 20% more similar to those of mammalian primates than those of birds. This helps reject the assumption that common physical characteristics between species is all that is needed to determine relatedness. 

Divergent evolution that produces the differences observed in the forest-dwelling, arboreal spider monkey and the nocturnal, airborne ghost bat can be reconciled through homology. Homologous characteristics are anatomical traits that are similar in two or more different species. For instance, the bone structure of a spider monkey’s wrist and fingers greatly resembles that of a bat’s wing or even a whale’s fin. These similarities are reinforced by phylogenetic evidence that supports the idea that physically dissimilar species can be evolutionarily related through anatomical and genetic similarities.

Phylogenetics traditionally investigates which molecular structures?

Adapted from Taking a Second Look: An Analysis of Genetic Markers in Species Relatedness by Joseph Ritchie (2014)

Phylogenetics is the study of genetic composition in various species and is used by evolutionary biologists to investigate similarities in the molecular sequences of proteins in varying organisms. The amino acid sequences that build proteins are used to construct mathmatical matrices that aid in determining evolutionary ties through the investigation of percentage similarities. The study of these matrices helps to expose evolutionary relationships between species that may not have the same existential characteristics.

Species adapt and evolve based on the pressures that exist in their environment. Climate, food source, and habitat availability are only a few factors that act on species adaptation. These stressors can alter the physical characteristics of organisms. This divergence in evolution has made it difficult to determine the interrelatedness of organisms by analyzing their physical characteristics alone.

For instance, looking only at physical characteristics, the ghost bat resembles a pigeon more than a spider monkey; however, phylogenetics has found that the amino acid sequences that construct the beta hemoglobin molecules of bats are 20% more similar to those of mammalian primates than those of birds. This helps reject the assumption that common physical characteristics between species is all that is needed to determine relatedness. 

Divergent evolution that produces the differences observed in the forest-dwelling, arboreal spider monkey and the nocturnal, airborne ghost bat can be reconciled through homology. Homologous characteristics are anatomical traits that are similar in two or more different species. For instance, the bone structure of a spider monkey’s wrist and fingers greatly resembles that of a bat’s wing or even a whale’s fin. These similarities are reinforced by phylogenetic evidence that supports the idea that physically dissimilar species can be evolutionarily related through anatomical and genetic similarities.

What mathematical tool is used in phylogenetics to study species interrelatedness?

Adapted from A Practical Treatise on the Hive and Honey-Bee by Lorenzo Lorraine Langstroth (1857 ed.)

Of all the numerous enemies of the honey-bee, the Bee-Moth (Tinea mellonella), in climates of hot summers, is by far the most to be dreaded. So widespread and fatal have been its ravages in this country that thousands have abandoned the cultivation of bees in despair, and in districts which once produced abundant supplies of the purest honey, bee-keeping has gradually dwindled down into a very insignificant pursuit. Contrivances almost without number have been devised to defend the bees against this invidious foe, but still it continues its desolating inroads, almost unchecked, laughing as it were to scorn at all the so-called "moth-proof" hives, and turning many of the ingenious fixtures designed to entrap or exclude it into actual aids and comforts in its nefarious designs.

I should feel but little confidence in being able to reinstate bee-keeping in our country into a certain and profitable pursuit if I could not show the apiarian in what way he can safely bid defiance to the pestiferous assaults of this, his most implacable enemy. I have patiently studied its habits for years, and I am at length able to announce a system of management founded upon the peculiar construction of my hives, which will enable the careful bee-keeper to protect his colonies against the monster. The bee-moth infects our apiaries, just as weeds take possession of a fertile soil. Before explaining the means upon which I rely to circumvent the moth, I will first give a brief description of its habits.

Swammerdam, towards the close of the seventeenth century, gave a very accurate description of this insect, which was then called by the very expressive name of the "bee-wolf." He has furnished good drawings of it, in all its changes, from the worm to the perfect moth, together with the peculiar webs or galleries that it constructs and from which the name of Tinea galleria or “gallery moth” has been given to it by some entomologists. He failed, however, to discriminate between the male and female, which, because they differ so much in size and appearance, he supposed to be two different species of the wax-moth. It seems to have been a great pest in his time, and even Virgil speaks of the "dirum tineæ genus," the dreadful offspring of the moth; that is the worm.

This destroyer usually makes its appearance about the hives in April or May, the time of its coming depending upon the warmth of the climate or the forwardness of the season. It is seldom seen on the wing (unless startled from its lurking place about the hive) until towards dark, and is evidently chiefly nocturnal in its habits. In dark cloudy days, however, I have noticed it on the wing long before sunset, and if several such days follow in succession, the female, oppressed with the urgent necessity of laying her eggs, may be seen endeavoring to gain admission to the hives. The female is much larger than the male, and "her color is deeper and more inclining to a darkish gray, with small spots or blackish streaks on the interior edge of her upper wings." The color of the male inclines more to a light gray; they might easily be mistaken for different species of moths. These insects are surprisingly agile, both on foot and on the wing. The motions of a bee are very slow in comparison. "They are," says Reaumur, "the most nimble-footed creatures that I know." "If the approach to the apiary be observed of a moonlight evening, the moths will be found flying or running round the hives, watching an opportunity to enter, whilst the bees that have to guard the entrances against their intrusion will be seen acting as vigilant sentinels, performing continual rounds near this important post, extending their antenna to the utmost, and moving them to the right and left alternately. Woe to the unfortunate moth that comes within their reach!" "It is curious," says Huber, "to observe how artfully the moth knows how to profit, to the disadvantage of the bees, which require much light for seeing objects; and the precautions taken by the latter in reconnoitering and expelling so dangerous an enemy."

The first paragraph establishes all of the following EXCEPT __________.

Adapted from A Practical Treatise on the Hive and Honey-Bee by Lorenzo Lorraine Langstroth (1857 ed.)

Of all the numerous enemies of the honey-bee, the Bee-Moth (Tinea mellonella), in climates of hot summers, is by far the most to be dreaded. So widespread and fatal have been its ravages in this country that thousands have abandoned the cultivation of bees in despair, and in districts which once produced abundant supplies of the purest honey, bee-keeping has gradually dwindled down into a very insignificant pursuit. Contrivances almost without number have been devised to defend the bees against this invidious foe, but still it continues its desolating inroads, almost unchecked, laughing as it were to scorn at all the so-called "moth-proof" hives, and turning many of the ingenious fixtures designed to entrap or exclude it into actual aids and comforts in its nefarious designs.

I should feel but little confidence in being able to reinstate bee-keeping in our country into a certain and profitable pursuit if I could not show the apiarian in what way he can safely bid defiance to the pestiferous assaults of this, his most implacable enemy. I have patiently studied its habits for years, and I am at length able to announce a system of management founded upon the peculiar construction of my hives, which will enable the careful bee-keeper to protect his colonies against the monster. The bee-moth infects our apiaries, just as weeds take possession of a fertile soil. Before explaining the means upon which I rely to circumvent the moth, I will first give a brief description of its habits.

Swammerdam, towards the close of the seventeenth century, gave a very accurate description of this insect, which was then called by the very expressive name of the "bee-wolf." He has furnished good drawings of it, in all its changes, from the worm to the perfect moth, together with the peculiar webs or galleries that it constructs and from which the name of Tinea galleria or “gallery moth” has been given to it by some entomologists. He failed, however, to discriminate between the male and female, which, because they differ so much in size and appearance, he supposed to be two different species of the wax-moth. It seems to have been a great pest in his time, and even Virgil speaks of the "dirum tineæ genus," the dreadful offspring of the moth; that is the worm.

This destroyer usually makes its appearance about the hives in April or May, the time of its coming depending upon the warmth of the climate or the forwardness of the season. It is seldom seen on the wing (unless startled from its lurking place about the hive) until towards dark, and is evidently chiefly nocturnal in its habits. In dark cloudy days, however, I have noticed it on the wing long before sunset, and if several such days follow in succession, the female, oppressed with the urgent necessity of laying her eggs, may be seen endeavoring to gain admission to the hives. The female is much larger than the male, and "her color is deeper and more inclining to a darkish gray, with small spots or blackish streaks on the interior edge of her upper wings." The color of the male inclines more to a light gray; they might easily be mistaken for different species of moths. These insects are surprisingly agile, both on foot and on the wing. The motions of a bee are very slow in comparison. "They are," says Reaumur, "the most nimble-footed creatures that I know." "If the approach to the apiary be observed of a moonlight evening, the moths will be found flying or running round the hives, watching an opportunity to enter, whilst the bees that have to guard the entrances against their intrusion will be seen acting as vigilant sentinels, performing continual rounds near this important post, extending their antenna to the utmost, and moving them to the right and left alternately. Woe to the unfortunate moth that comes within their reach!" "It is curious," says Huber, "to observe how artfully the moth knows how to profit, to the disadvantage of the bees, which require much light for seeing objects; and the precautions taken by the latter in reconnoitering and expelling so dangerous an enemy."

Which of the following statements about bees is supported by the passage?