Adapted from “Introduced Species That Have Become Pests” in Our Vanishing Wild Life, Its Extermination and Protection by William Temple Hornaday (1913)

The man who successfully transplants or "introduces" into a new habitat any persistent species of living thing assumes a very grave responsibility. Every introduced species is doubtful gravel until panned out. The enormous losses that have been inflicted upon the world through the perpetuation of follies with wild vertebrates and insects would, if added together, be enough to purchase a principality. The most aggravating feature of these follies in transplantation is that never yet have they been made severely punishable. We are just as careless and easygoing on this point as we were about the government of the Yellowstone Park in the days when Howell and other poachers destroyed our first national bison herd, and when caught red-handed—as Howell was, skinning seven Park bison cows—could not be punished for it, because there was no penalty prescribed by any law. Today, there is a way in which any revengeful person could inflict enormous damage on the entire South, at no cost to himself, involve those states in enormous losses and the expenditure of vast sums of money, yet go absolutely unpunished!

The gypsy moth is a case in point. This winged calamity was imported at Maiden, Massachusetts, near Boston, by a French entomologist, Mr. Leopold Trouvelot, in 1868 or 69. History records the fact that the man of science did not purposely set free the pest. He was endeavoring with live specimens to find a moth that would produce a cocoon of commercial value to America, and a sudden gust of wind blew out of his study, through an open window, his living and breeding specimens of the gypsy moth. The moth itself is not bad to look at, but its larvae is a great, overgrown brute with an appetite like a hog. Immediately Mr. Trouvelot sought to recover his specimens, and when he failed to find them all, like a man of real honor, he notified the State authorities of the accident. Every effort was made to recover all the specimens, but enough escaped to produce progeny that soon became a scourge to the trees of Massachusetts. The method of the big, nasty-looking mottled-brown caterpillar was very simple. It devoured the entire foliage of every tree that grew in its sphere of influence.

The gypsy moth spread with alarming rapidity and persistence. In course of time, the state authorities of Massachusetts were forced to begin a relentless war upon it, by poisonous sprays and by fire. It was awful! Up to this date (1912) the New England states and the United States Government service have expended in fighting this pest about $7,680,000!

The spread of this pest has been retarded, but the gypsy moth never will be wholly stamped out. Today it exists in Rhode Island, Connecticut, and New Hampshire, and it is due to reach New York at an early date. It is steadily spreading in three directions from Boston, its original point of departure, and when it strikes the State of New York, we, too, will begin to pay dearly for the Trouvelot experiment.

Which of the following best paraphrases the underlined sentence, “Every introduced species is doubtful gravel until panned out”?

Adapted from “Feathers of Sea Birds and Wild Fowl for Bedding” from The Utility of Birds by Edward Forbush (ed. 1922)

In the colder countries of the world, the feathers and down of waterfowl have been in great demand for centuries as filling for beds and pillows. Such feathers are perfect non-conductors of heat, and beds, pillows, or coverlets filled with them represent the acme of comfort and durability. The early settlers of New England saved for such purposes the feathers and down from the thousands of wild-fowl which they killed, but as the population increased in numbers, the quantity thus furnished was insufficient, and the people sought a larger supply in the vast colonies of ducks and geese along the Labrador coast. 

The manner in which the feathers and down were obtained, unlike the method practiced in Iceland, did not tend to conserve and protect the source of supply. In Iceland, the people have continued to receive for many years a considerable income by collecting eider down, but there they do not “kill the goose that lays the golden eggs.” Ducks line their nests with down plucked from their own breasts and that of the eider is particularly valuable for bedding. In Iceland, these birds are so carefully protected that they have become as tame and unsuspicious as domestic fowls In North America. Where they are constantly hunted they often conceal their nests in the midst of weeds or bushes, but in Iceland, they make their nests and deposit their eggs in holes dug for them in the sod. A supply of the ducks is maintained so that the people derive from them an annual income.

In North America, quite a different policy was pursued. The demand for feathers became so great in the New England colonies about the middle of the eighteenth century that vessels were fitted out there for the coast of Labrador for the express purpose of securing the feathers and down of wild fowl. Eider down having become valuable and these ducks being in the habit of congregating by thousands on barren islands of the Labrador coast, the birds became the victims of the ships’ crews. As the ducks molt all their primary feathers at once in July or August and are then quite incapable of flight and the young birds are unable to fly until well grown, the hunters were able to surround the helpless birds, drive them together, and kill them with clubs. Otis says that millions of wildfowl were thus destroyed and that in a few years their haunts were so broken up by this wholesale slaughter and their numbers were so diminished that feather voyages became unprofitable and were given up. 

This practice, followed by the almost continual egging, clubbing, shooting, etc. by Labrador fishermen, may have been a chief factor in the extinction of the Labrador duck, that species of supposed restricted breeding range. No doubt had the eider duck been restricted in its breeding range to the islands of Labrador, it also would have been exterminated long ago.

Which of the following best restates the meaning of the underlined phrase “as the population increased in numbers, the quantity thus furnished was insufficient”?

Adapted from “Humming-Birds: As Illustrating the Luxuriance of Tropical Nature” in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The food of hummingbirds has been a matter of much controversy. All the early writers down to Buffon believed that they lived solely on the nectar of flowers, but since that time, every close observer of their habits maintains that they feed largely, and in some cases wholly, on insects. Azara observed them on the La Plata in winter taking insects out of the webs of spiders at a time and place where there were no flowers. Bullock, in Mexico, declares that he saw them catch small butterflies, and that he found many kinds of insects in their stomachs. Waterton made a similar statement. Hundreds and perhaps thousands of specimens have since been dissected by collecting naturalists, and in almost every instance their stomachs have been found full of insects, sometimes, but not generally, mixed with a proportion of honey. Many of them in fact may be seen catching gnats and other small insects just like fly-catchers, sitting on a dead twig over water, darting off for a time in the air, and then returning to the twig. Others come out just at dusk, and remain on the wing, now stationary, now darting about with the greatest rapidity, imitating in a limited space the evolutions of the goatsuckers, and evidently for the same end and purpose. Mr. Gosse also remarks, ” All the hummingbirds have more or less the habit, when in flight, of pausing in the air and throwing the body and tail into rapid and odd contortions. This is most observable in the Polytmus, from the effect that such motions have on the long feathers of the tail. That the object of these quick turns is the capture of insects, I am sure, having watched one thus engaged pretty close to me.”

What can we infer from the author’s use of the underlined phrase, “sometimes, but not generally”?

Adapted from “Humming-Birds: As Illustrating the Luxuriance of Tropical Nature” in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The food of hummingbirds has been a matter of much controversy. All the early writers down to Buffon believed that they lived solely on the nectar of flowers, but since that time, every close observer of their habits maintains that they feed largely, and in some cases wholly, on insects. Azara observed them on the La Plata in winter taking insects out of the webs of spiders at a time and place where there were no flowers. Bullock, in Mexico, declares that he saw them catch small butterflies, and that he found many kinds of insects in their stomachs. Waterton made a similar statement. Hundreds and perhaps thousands of specimens have since been dissected by collecting naturalists, and in almost every instance their stomachs have been found full of insects, sometimes, but not generally, mixed with a proportion of honey. Many of them in fact may be seen catching gnats and other small insects just like fly-catchers, sitting on a dead twig over water, darting off for a time in the air, and then returning to the twig. Others come out just at dusk, and remain on the wing, now stationary, now darting about with the greatest rapidity, imitating in a limited space the evolutions of the goatsuckers, and evidently for the same end and purpose. Mr. Gosse also remarks, ” All the hummingbirds have more or less the habit, when in flight, of pausing in the air and throwing the body and tail into rapid and odd contortions. This is most observable in the Polytmus, from the effect that such motions have on the long feathers of the tail. That the object of these quick turns is the capture of insects, I am sure, having watched one thus engaged pretty close to me.”

The meaning of the underlined phrase “on the wing” is __________.

Adapted from “Comets” by Camille Flammarion in Wonders of Earth, Sea, and Sky (1902, ed. Edward Singleton Holden)

The history of a comet would be an instructive episode of the great history of the heavens. In it could be brought together the description of the progressive movement of human thought, as well as the astronomical theory of these extraordinary bodies. Let us take, for example, one of the most memorable and best-known comets, and give an outline of its successive passages near the Earth. Like the planetary worlds, comets belong to the solar system, and are subject to the rule of the Star King. It is the universal law of gravitation which guides their path; solar attraction governs them, as it governs the movement of the planets and the small satellites. The chief point of difference between them and the planets is that their orbits are very elongated, and instead of being nearly circular, they take the elliptical form. In consequence of the nature of these orbits, the same comet may approach very near the sun, and afterwards travel from it to immense distances.

Thus, the period of the Comet of 1680 has been estimated at three thousand years. It approaches the sun, so as to be nearer to it than our moon is to us, whilst it recedes to a distance 853 times greater than the distance of the Earth from the sun. On the 17th of December, 1680, it was at its perihelion—that is, at its greatest proximity to the sun; it is now continuing its path beyond the Neptunian orbit. Its velocity varies according to its distance from the solar body. At its perihelion it travels thousands of leagues per minute; at its aphelion it does not pass over more than a few yards.  

Its proximity to the Sun in its passage near that body caused Newton to think that it received a heat twenty-eight thousand times greater than that we experience at the summer solstice, and that this heat being two thousand times greater than that of red-hot iron, an iron globe of the same dimensions would be fifty thousand years entirely losing its heat. Newton added that in the end, comets will approach so near the sun that they will not be able to escape the preponderance of its attraction, and that they will fall one after the other into this brilliant body, thus keeping up the heat which it perpetually pours out into space. Such is the deplorable end assigned to comets by the author of the Principia, an end which makes De la Brétonne say to Rétif: "An immense comet, already larger than Jupiter, was again increased in its path by being blended with six other dying comets. Thus displaced from its ordinary route by these slight shocks, it did not pursue its true elliptical orbit; so that the unfortunate thing was precipitated into the devouring centre of the Sun." "It is said," added he, "that the poor comet, thus burned alive, sent forth dreadful cries!"

What does the author most nearly mean when he says “Like the planetary worlds, comets belong to the solar system, and are subject to the rule of the Star King.”?

Adapted from "The Man-Like Apes" by T. H. Huxley in A Book of Natural History (1902, ed. David Starr Jordan)

The orangutan is found only in Sumatra and Borneo, and is common in either of these islands—in both of which it occurs always in low, flat plains, never in the mountains. It loves the densest and most sombre of the forests, which extend from the seashore inland, and thus is found only in the eastern half of Sumatra, where alone such forests occur, though, occasionally, it strays over to the western side. On the other hand, it is generally distributed through Borneo, except in the mountains, or where the population is dense. In favorable places the hunter may, by good fortune, see three or four in a day.

Except in the pairing time, the old males usually live by themselves. The old females and the immature males, on the other hand, are often met with in twos and threes, and the former occasionally have young with them, though the pregnant females usually separate themselves, and sometimes remain apart after they have given birth to their offspring. The young orangs seem to remain unusually long under their mother’s protection, probably in consequence of their slow growth. While climbing, the mother always carries her young against her bosom, the young holding on by the mother’s hair. At what time of life the orangutan becomes capable of propagation, and how long the females go with young is unknown, but it is probable that they are not adult until they arrive at ten or fifteen years of age. A female which lived for five years at Batavia had not attained one-third the height of the wild females. It is probable that, after reaching adult years, they go on growing, though slowly, and that they live to forty or fifty years. The Dyaks tell of old orangs that have not only lost all their teeth, but which find it so troublesome to climb that they maintain themselves on windfalls and juicy herbage.

What does the author most nearly mean when he says “On the other hand, it is generally distributed through Borneo, except in the mountains, or where the population is dense”?

Adapted from Volume Four of The Natural History of Animals: The Animal Life of the World in Its Various Aspects and Relations by James Richard Ainsworth Davis (1903)

The examples of protective resemblance so far quoted are mostly permanent adaptations to one particular sort of surrounding. There are, however, numerous animals which possess the power of adjusting their color more or less rapidly so as to harmonize with a changing environment.

Some of the best known of these cases are found among those mammals and birds that inhabit countries more or less covered with snow during a part of the year. A good instance is afforded by the Irish or variable hare, which is chiefly found in Ireland and Scotland. In summer, this looks very much like an ordinary hare, though rather grayer in tint and smaller in size, but in winter it becomes white with the exception of the black tips to the ears. Investigations that have been made on the closely allied American hare seem to show that the phenomenon is due to the growth of new hairs of white hue. 

The common stoat is subject to similar color change in the northern parts of its range. In summer it is of a bright reddish brown color with the exception of the under parts, which are yellowish white, and the end of the tail, which is black. But in winter, the entire coat, save only the tip of the tail, becomes white, and in that condition the animal is known as an ermine. A similar example is afforded by the weasel. The seasonal change in the vegetarian Irish hare is purely of protective character, but in such an actively carnivorous creature as a stoat or weasel, it is aggressive as well, rendering the animal inconspicuous to its prey.

The phrase “harmonize with,” underlined in the first paragraph, most closely means __________.

Adapted from On the Origin of Species by Charles Darwin (1859)

How will the struggle for existence, discussed too briefly in the last chapter, act in regard to variation? Can the principle of selection, which we have seen is so potent in the hands of man, apply in nature? I think we shall see that it can act most effectually. Let it be borne in mind in what an endless number of strange peculiarities our domestic productions, and, in a lesser degree, those under nature, vary; and how strong the hereditary tendency is. Under domestication, it may be truly said that the whole organization becomes in some degree plastic. Let it be borne in mind how infinitely complex and close-fitting are the mutual relations of all organic beings to each other and to their physical conditions of life. Can it, then, be thought improbable, seeing that variations useful to man have undoubtedly occurred, that other variations useful in some way to each being in the great and complex battle of life, should sometimes occur in the course of thousands of generations? If such do occur, can we doubt (remembering that many more individuals are born than can possibly survive) that individuals having any advantage, however slight, over others, would have the best chance of surviving and of procreating their kind? On the other hand, we may feel sure that any variation in the least degree injurious would be rigidly destroyed. This preservation of favorable variations and the rejection of injurious variations, I call Natural Selection. Variations neither useful nor injurious would not be affected by natural selection, and would be left a fluctuating element, as perhaps we see in the species called polymorphic.

We shall best understand the probable course of natural selection by taking the case of a country undergoing some physical change, for instance, of climate. The proportional numbers of its inhabitants would almost immediately undergo a change, and some species might become extinct. We may conclude, from what we have seen of the intimate and complex manner in which the inhabitants of each country are bound together, that any change in the numerical proportions of some of the inhabitants, independently of the change of climate itself, would most seriously affect many of the others. If the country were open on its borders, new forms would certainly immigrate, and this also would seriously disturb the relations of some of the former inhabitants. Let it be remembered how powerful the influence of a single introduced tree or mammal has been shown to be. But in the case of an island, or of a country partly surrounded by barriers, into which new and better adapted forms could not freely enter, we should then have places in the economy of nature which would assuredly be better filled up, if some of the original inhabitants were in some manner modified; for, had the area been open to immigration, these same places would have been seized on by intruders. In such case, every slight modification, which in the course of ages chanced to arise, and which in any way favoured the individuals of any of the species, by better adapting them to their altered conditions, would tend to be preserved; and natural selection would thus have free scope for the work of improvement.

Darwin is most likely a __________.

Adapted from On the Origin of Species by Charles Darwin (1859)

How will the struggle for existence, discussed too briefly in the last chapter, act in regard to variation? Can the principle of selection, which we have seen is so potent in the hands of man, apply in nature? I think we shall see that it can act most effectually. Let it be borne in mind in what an endless number of strange peculiarities our domestic productions, and, in a lesser degree, those under nature, vary; and how strong the hereditary tendency is. Under domestication, it may be truly said that the whole organization becomes in some degree plastic. Let it be borne in mind how infinitely complex and close-fitting are the mutual relations of all organic beings to each other and to their physical conditions of life. Can it, then, be thought improbable, seeing that variations useful to man have undoubtedly occurred, that other variations useful in some way to each being in the great and complex battle of life, should sometimes occur in the course of thousands of generations? If such do occur, can we doubt (remembering that many more individuals are born than can possibly survive) that individuals having any advantage, however slight, over others, would have the best chance of surviving and of procreating their kind? On the other hand, we may feel sure that any variation in the least degree injurious would be rigidly destroyed. This preservation of favorable variations and the rejection of injurious variations, I call Natural Selection. Variations neither useful nor injurious would not be affected by natural selection, and would be left a fluctuating element, as perhaps we see in the species called polymorphic.

We shall best understand the probable course of natural selection by taking the case of a country undergoing some physical change, for instance, of climate. The proportional numbers of its inhabitants would almost immediately undergo a change, and some species might become extinct. We may conclude, from what we have seen of the intimate and complex manner in which the inhabitants of each country are bound together, that any change in the numerical proportions of some of the inhabitants, independently of the change of climate itself, would most seriously affect many of the others. If the country were open on its borders, new forms would certainly immigrate, and this also would seriously disturb the relations of some of the former inhabitants. Let it be remembered how powerful the influence of a single introduced tree or mammal has been shown to be. But in the case of an island, or of a country partly surrounded by barriers, into which new and better adapted forms could not freely enter, we should then have places in the economy of nature which would assuredly be better filled up, if some of the original inhabitants were in some manner modified; for, had the area been open to immigration, these same places would have been seized on by intruders. In such case, every slight modification, which in the course of ages chanced to arise, and which in any way favoured the individuals of any of the species, by better adapting them to their altered conditions, would tend to be preserved; and natural selection would thus have free scope for the work of improvement.

From this passage we can infer that Darwin __________.

Adapted from "Recent Views as to Direct Action of Light on the Colors of Flowers and Fruits" in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The theory that the brilliant colors of flowers and fruits is due to the direct action of light has been supported by a recent writer by examples taken from the arctic instead of from the tropical flora. In the arctic regions, vegetation is excessively rapid during the short summer, and this is held to be due to the continuous action of light throughout the long summer days. “The further we advance towards the north, the more the leaves of plants increase in size as if to absorb a greater proportion of the solar rays. M. Grisebach says that during a journey in Norway he observed that the majority of deciduous trees had already, at the 60th degree of latitude, larger leaves than in Germany, while M. Ch. Martins has made a similar observation as regards the leguminous plants cultivated in Lapland.” The same writer goes on to say that all the seeds of cultivated plants acquire a deeper color the further north they are grown, white haricots becoming brown or black, and white wheat becoming brown, while the green color of all vegetation becomes more intense. The flowers also are similarly changed: those which are white or yellow in central Europe becoming red or orange in Norway. This is what occurs in the Alpine flora, and the cause is said to be the same in both—the greater intensity of the sunlight. In the one the light is more persistent, in the other more intense because it traverses a less thickness of atmosphere.

Admitting the facts as above stated to be in themselves correct, they do not by any means establish the theory founded on them; and it is curious that Grisebach, who has been quoted by this writer for the fact of the increased size of the foliage, gives a totally different explanation of the more vivid colors of Arctic flowers. He says, “We see flowers become larger and more richly colored in proportion as, by the increasing length of winter, insects become rarer, and their cooperation in the act of fecundation is exposed to more uncertain chances.” (Vegetation du Globe, col. i. p. 61—French translation.) This is the theory here adopted to explain the colors of Alpine plants, and we believe there are many facts that will show it to be the preferable one. The statement that the white and yellow flowers of temperate Europe become red or golden in the Arctic regions must we think be incorrect. By roughly tabulating the colors of the plants given by Sir Joseph Hooker as permanently Arctic, we find among fifty species with more or less conspicuous flowers, twenty-five white, twelve yellow, eight purple or blue, three lilac, and two red or pink; showing a very similar proportion of white and yellow flowers to what obtains further south.

The author of the passage is most likely __________.