Passage adapted from The Extermination of the American Bison, by William Hornaday (1889).

The history of the buffalo’s daily life and habits should begin with the “running season.” This period occupied the months of August and September, and was characterized by a degree of excitement and activity throughout the entire herd quite foreign to the ease-loving and even slothful nature which was so noticeable a feature of the bison’s character at all other times.

The mating season occurred when the herd was on its summer range. The spring calves were from two to four months old. Through continued feasting on the new crop of buffalo-grass and bunch-grass—the most nutritious in the world, perhaps—every buffalo in the herd had grown round-sided, fat, and vigorous. The faded and weather-beaten suit of winter hair had by that time fallen off and given place to the new coat of dark gray and black, and, excepting for the shortness of his hair, the buffalo was in prime condition.

During the “running season,” as it was called by the plainsmen, the whole nature of the herd was completely changed. Instead of being broken up into countless small groups and dispersed over a vast extent of territory, the herd came together in a dense and confused mass of many thousand individuals, so closely congregated as to actually blacken the face of the landscape. As if by a general and irresistible impulse, every straggler would be drawn to the common center, and for miles on every side of the great herd the country would be found entirely deserted.

At this time the herd itself became a seething mass of activity and excitement. As usual under such conditions, the bulls were half the time chasing the cows, and fighting each other during the other half. These actual combats, which were always of short duration and over in a few seconds after the actual collision took place, were preceded by the usual threatening demonstrations, in which the bull lowers his head until his nose almost touches the ground, roars like a fog-horn until the earth seems to fairly tremble with the vibration, glares madly upon his adversary with half-white eyeballs, and with his forefeet paws up the dry earth and throws it upward in a great cloud of dust high above his back. At such times the mingled roaring—it can not truthfully be described as lowing or bellowing—of a number of huge bulls unite and form a great volume of sound like distant thunder, which has often been heard at a distance of from 1 to 3 miles. I have even been assured by old plainsmen that under favorable atmospheric conditions such sounds have been heard five miles.

It can be inferred by the passage that the plainsmen are __________.

Adapted from "Taking a Second Look: An Analysis of Genetic Markers in Species Relatedness" by Joseph Ritchie (2014)

Phylogenetics is the study of genetic composition in various species and is used by evolutionary biologists to investigate similarities in the molecular sequences of proteins in varying organisms. The amino acid sequences that build proteins are used to construct mathematical matrices that aid in determining evolutionary ties through the investigation of percentage similarities. The study of these matrices helps to expose evolutionary relationships between species that may not have the same overt characteristics.

Species adapt and evolve based on the pressures that exist in their environment. Climate, food source, and habitat availability are only a few factors that act on species adaptation. These stressors can alter the physical characteristics of organisms. This divergence in evolution has made it difficult to determine the interrelatedness of organisms by analyzing their physical characteristics alone.

For instance, looking only at physical characteristics, the ghost bat resembles a pigeon more than a spider monkey; however, phylogenetics has found that the amino acid sequences that construct the beta hemoglobin molecules of bats are twenty percent more similar to those of mammalian primates than those of birds. This helps reject the assumption that common physical characteristics between species are all that is needed to determine relatedness. 

The differences produced by divergent evolution observed in the forest-dwelling, arboreal spider monkey and the nocturnal, airborne ghost bat can be reconciled through homology. Homologous characteristics are anatomical traits that are similar in two or more different species. For instance, the bone structure of a spider monkey’s wrist and fingers greatly resembles that of a bat’s wing or even a whale’s fin. These similarities are reinforced by phylogenetic evidence that supports the idea that physically dissimilar species can be evolutionarily related through anatomical and genetic similarities.

A scientist studied the relatedness of several reptilian species solely by investigating fossil evidence and has concluded that physical characteristics alone are enough to determine species relatedness. Would this scientist agree with the claims made by phylogenetic research?

Adapted from "Taking a Second Look: An Analysis of Genetic Markers in Species Relatedness" by Joseph Ritchie (2014)

Phylogenetics is the study of genetic composition in various species and is used by evolutionary biologists to investigate similarities in the molecular sequences of proteins in varying organisms. The amino acid sequences that build proteins are used to construct mathematical matrices that aid in determining evolutionary ties through the investigation of percentage similarities. The study of these matrices helps to expose evolutionary relationships between species that may not have the same overt characteristics.

Species adapt and evolve based on the pressures that exist in their environment. Climate, food source, and habitat availability are only a few factors that act on species adaptation. These stressors can alter the physical characteristics of organisms. This divergence in evolution has made it difficult to determine the interrelatedness of organisms by analyzing their physical characteristics alone.

For instance, looking only at physical characteristics, the ghost bat resembles a pigeon more than a spider monkey; however, phylogenetics has found that the amino acid sequences that construct the beta hemoglobin molecules of bats are twenty percent more similar to those of mammalian primates than those of birds. This helps reject the assumption that common physical characteristics between species are all that is needed to determine relatedness. 

The differences produced by divergent evolution observed in the forest-dwelling, arboreal spider monkey and the nocturnal, airborne ghost bat can be reconciled through homology. Homologous characteristics are anatomical traits that are similar in two or more different species. For instance, the bone structure of a spider monkey’s wrist and fingers greatly resembles that of a bat’s wing or even a whale’s fin. These similarities are reinforced by phylogenetic evidence that supports the idea that physically dissimilar species can be evolutionarily related through anatomical and genetic similarities.

The anatomical similarities between a horse's legs and a seal's flippers is best explained by which of the following?

Adapted from The Effects of Cross & Self-Fertilisation in the Vegetable Kingdom by Charles Darwin (1876)

As it is impossible to exclude such minute pollen-carrying insects as Thrips, flowers which it was intended to fertilise with their own pollen may sometimes have been afterwards crossed with pollen brought by these insects from another flower on the same plant; but as we shall hereafter see, a cross of this kind does not produce any effect, or at most only a slight one. When two or more plants were placed near one another under the same net, as was often done, there is some real though not great danger of the flowers which were believed to be self-fertilised being afterwards crossed with pollen brought by Thrips from a distinct plant. I have said that the danger is not great because I have often found that plants which are self-sterile, unless aided by insects, remained sterile when several plants of the same species were placed under the same net. If, however, the flowers which had been presumably self-fertilised by me were in any case afterwards crossed by Thrips with pollen brought from a distinct plant, crossed seedlings would have been included amongst the self-fertilised; but it should be especially observed that this occurrence would tend to diminish and not to increase any superiority in average height, fertility, etc., of the crossed over the self-fertilised plants.

As the flowers which were crossed were never castrated, it is probable or even almost certain that I sometimes failed to cross-fertilise them effectually, and that they were afterwards spontaneously self-fertilised. This would have been most likely to occur with dichogamous species, for without much care it is not easy to perceive whether their stigmas are ready to be fertilised when the anthers open. But in all cases, as the flowers were protected from wind, rain, and the access of insects, any pollen placed by me on the stigmatic surface whilst it was immature, would generally have remained there until the stigma was mature; and the flowers would then have been crossed as was intended. Nevertheless, it is highly probable that self-fertilised seedlings have sometimes by this means got included amongst the crossed seedlings. The effect would be, as in the former case, not to exaggerate but to diminish any average superiority of the crossed over the self-fertilised plants.

Errors arising from the two causes just named, and from others,—such as some of the seeds not having been thoroughly ripened, though care was taken to avoid this error—the sickness or unperceived injury of any of the plants,—will have been to a large extent eliminated, in those cases in which many crossed and self-fertilised plants were measured and an average struck. Some of these causes of error will also have been eliminated by the seeds having been allowed to germinate on bare damp sand, and being planted in pairs; for it is not likely that ill-matured and well-matured, or diseased and healthy seeds, would germinate at exactly the same time. The same result will have been gained in the several cases in which only a few of the tallest, finest, and healthiest plants on each side of the pots were measured.

Kolreuter and Gartner have proved that with some plants several, even as many as from fifty to sixty, pollen-grains are necessary for the fertilisation of all the ovules in the ovarium. Naudin also found in the case of Mirabilis that if only one or two of its very large pollen-grains were placed on the stigma, the plants raised from such seeds were dwarfed. I was therefore careful to give an amply sufficient supply of pollen, and generally covered the stigma with it; but I did not take any special pains to place exactly the same amount on the stigmas of the self-fertilised and crossed flowers. After having acted in this manner during two seasons, I remembered that Gartner thought, though without any direct evidence, that an excess of pollen was perhaps injurious. It was therefore necessary to ascertain whether the fertility of the flowers was affected by applying a rather small and an extremely large quantity of pollen to the stigma. Accordingly a very small mass of pollen-grains was placed on one side of the large stigma in sixty-four flowers of Ipomoea purpurea, and a great mass of pollen over the whole surface of the stigma in sixty-four other flowers. In order to vary the experiment, half the flowers of both lots were on plants produced from self-fertilised seeds, and the other half on plants from crossed seeds. The sixty-four flowers with an excess of pollen yielded sixty-one capsules; and excluding four capsules, each of which contained only a single poor seed, the remainder contained on an average 5.07 seeds per capsule. The sixty-four flowers with only a little pollen placed on one side of the stigma yielded sixty-three capsules, and excluding one from the same cause as before, the remainder contained on an average 5.129 seeds. So that the flowers fertilised with little pollen yielded rather more capsules and seeds than did those fertilised with an excess; but the difference is too slight to be of any significance. On the other hand, the seeds produced by the flowers with an excess of pollen were a little heavier of the two; for 170 of them weighed 79.67 grains, whilst 170 seeds from the flowers with very little pollen weighed 79.20 grains. Both lots of seeds having been placed on damp sand presented no difference in their rate of germination. We may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.

Based on the passage, a purpose for the damp sand was to __________.

Adapted from Ice-Caves of France and Switzerland by George Forrest Browne (1865)

This account states that the cave is in the county of Thorn, among the lowest spurs of the Carpathians. The entrance, which faces the north, and is exposed to the cold winds from the snowy part of the Carpathian range, is eighteen fathoms high and nine broad; and the cave spreads out laterally, and descends to a point fifty fathoms below the entrance, where it is twenty-six fathoms in breadth, and of irregular height. Beyond this no one had at that time penetrated, on account of the unsafe footing, although many distant echoes were returned by the farther recesses of the cave; indeed, to get even so far as this, much step-cutting was necessary.

When the external frost of winter comes on, the account proceeds, the effect in the cave is the same as if fires had been lighted there: the ice melts, and swarms of flies and bats and hares take refuge in the interior from the severity of the winter. As soon as spring arrives, the warmth of winter disappears from the interior, water exudes from the roof and is converted into ice, while the more abundant supplies which pour down on to the sandy floor are speedily frozen there. In the dog-days, the frost is so intense that a small icicle becomes in one day a huge mass of ice; but a cool day promptly brings a thaw, and the cave is looked upon as a barometer, not merely feeling, but also presaging, the changes of weather. The people of the neighborhood, when employed in field-work, arrange their labour so that the mid-day meal may be taken near the cave, when they either ice the water they have brought with them, or drink the melted ice, which they consider very good for the stomach. It had been calculated that six hundred weekly carts would not be sufficient to keep the cavern free from ice. The ground above the cave is peculiarly rich in grass.

In explanation of these phenomena, Bell threw out the following suggestions, which need no comment. The earth being of itself cold and damp, the external heat of the atmosphere, by partially penetrating into the ground, drives in this native cold to the inner parts of the earth, and makes the cold there more dense. On the other hand, when the external air is cold, it draws forth towards the surface the heat there may be in the inner part of the earth, and thus makes caverns warm. In support and illustration of this view, he states that in the hotter parts of Hungary, when the people wish to cool their wine, they dig a hole two feet deep, and place in it the flagon of wine, and, after filling up the hole again, light a blazing fire upon the surface, which cools the wine as if the flagon had been laid in ice. He also suggests that possibly the cold winds from the Carpathians bring with them imperceptible particles of snow, which reach the water of the cave, and convert it into ice. Further, the rocks of the Carpathians abound in salts, niter, alum, etc., which may, perhaps, mingle with such snowy particles, and produce the ordinary effect of the snow and salt in the artificial production of ice.

In the third paragraph, the information about Bell's hypothesis serves to __________.

Adapted from A Practical Treatise on the Hive and Honey-Bee by Lorenzo Lorraine Langstroth (1857 ed.)

Of all the numerous enemies of the honey-bee, the Bee-Moth (Tinea mellonella), in climates of hot summers, is by far the most to be dreaded. So widespread and fatal have been its ravages in this country that thousands have abandoned the cultivation of bees in despair, and in districts which once produced abundant supplies of the purest honey, bee-keeping has gradually dwindled down into a very insignificant pursuit. Contrivances almost without number have been devised to defend the bees against this invidious foe, but still it continues its desolating inroads, almost unchecked, laughing as it were to scorn at all the so-called "moth-proof" hives, and turning many of the ingenious fixtures designed to entrap or exclude it into actual aids and comforts in its nefarious designs.

I should feel but little confidence in being able to reinstate bee-keeping in our country into a certain and profitable pursuit if I could not show the apiarian in what way he can safely bid defiance to the pestiferous assaults of this, his most implacable enemy. I have patiently studied its habits for years, and I am at length able to announce a system of management founded upon the peculiar construction of my hives, which will enable the careful bee-keeper to protect his colonies against the monster. The bee-moth infects our apiaries, just as weeds take possession of a fertile soil. Before explaining the means upon which I rely to circumvent the moth, I will first give a brief description of its habits.

Swammerdam, towards the close of the seventeenth century, gave a very accurate description of this insect, which was then called by the very expressive name of the "bee-wolf." He has furnished good drawings of it, in all its changes, from the worm to the perfect moth, together with the peculiar webs or galleries that it constructs and from which the name of Tinea galleria or “gallery moth” has been given to it by some entomologists. He failed, however, to discriminate between the male and female, which, because they differ so much in size and appearance, he supposed to be two different species of the wax-moth. It seems to have been a great pest in his time, and even Virgil speaks of the "dirum tineæ genus," the dreadful offspring of the moth; that is the worm.

This destroyer usually makes its appearance about the hives in April or May, the time of its coming depending upon the warmth of the climate or the forwardness of the season. It is seldom seen on the wing (unless startled from its lurking place about the hive) until towards dark, and is evidently chiefly nocturnal in its habits. In dark cloudy days, however, I have noticed it on the wing long before sunset, and if several such days follow in succession, the female, oppressed with the urgent necessity of laying her eggs, may be seen endeavoring to gain admission to the hives. The female is much larger than the male, and "her color is deeper and more inclining to a darkish gray, with small spots or blackish streaks on the interior edge of her upper wings." The color of the male inclines more to a light gray; they might easily be mistaken for different species of moths. These insects are surprisingly agile, both on foot and on the wing. The motions of a bee are very slow in comparison. "They are," says Reaumur, "the most nimble-footed creatures that I know." "If the approach to the apiary be observed of a moonlight evening, the moths will be found flying or running round the hives, watching an opportunity to enter, whilst the bees that have to guard the entrances against their intrusion will be seen acting as vigilant sentinels, performing continual rounds near this important post, extending their antenna to the utmost, and moving them to the right and left alternately. Woe to the unfortunate moth that comes within their reach!" "It is curious," says Huber, "to observe how artfully the moth knows how to profit, to the disadvantage of the bees, which require much light for seeing objects; and the precautions taken by the latter in reconnoitering and expelling so dangerous an enemy."

In the third paragraph the information about Swammerdam's name for the moth serves to __________.

"Interpreting the Copernican Revolution" by Matthew Minerd (2014)

The expressions of one discipline can often alter the way that other subjects understand themselves. Among such cases are numbered the investigations of Nicolaus Copernicus. Copernicus is best known for his views concerning heliocentrism, a view which eventually obliterated many aspects of the ancient/medieval worldview, at least from the standpoint of physical science. It had always been the natural view of mankind that the earth stood at the center of the universe, a fixed point in reference to the rest of the visible bodies. The sun, stars, and planets all rotated around the earth.

With time, this viewpoint became one of the major reference points for modern life. It provided a provocative image that was used—and often abused—by many people for various purposes. For those who wished to weaken the control of religion on mankind, it was said that the heliocentric outlook proved man’s insignificance. In contrast with earlier geocentrism, heliocentrism was said to show that man is not the center of the universe. He is merely one small being in the midst of a large cosmos. However, others wished to use the “Copernican Revolution” in a very different manner. These thinkers wanted to show that there was another “recentering” that had to happen. Once upon a time, we talked about the world. Now, however, it was necessary to talk of man as the central reference point. Just as the solar system was “centered” on the sun, so too should the sciences be centered on the human person.

However, both of these approaches are fraught with problems. Those who wished to undermine the religious mindset rather misunderstood the former outlook on the solar system. The earlier geocentric mindset did not believe that the earth was the most important body in the heavens. Instead, many ancient and medieval thinkers believed that the highest “sphere” above the earth was the most important being in the physical universe. Likewise, the so-called “Copernican Revolution” in physics was different from the one applied to the human person. Copernicus’ revolution showed that the human point of view was not the center, whereas the later forms of “Copernican revolution” wished to show just the opposite.

Of course, there are many complexities in the history of such important changes in scientific outlook. Nevertheless, it is fascinating to see the wide-reaching effects of such discoveries, even when they have numerous, ambiguous effects.

What can we say of the effects of Copernicus’ discoveries on the reigning medieval way of looking at the world?

"Darwinism's Effect on Science" by Matthew Minerd (2014)

For much of the history of human thought, the sciences have studied subjects that seemed to be eternal and unchanging. Even the basic laws of the Nile’s flooding were investigated in the hopes of finding never-altering laws. Similarly, the scientific investigations of the ancient Near East and Greece into the regular laws of the stars ultimately looked for constant patterns. This overall pattern of scientific reasoning has left deep marks on the minds of almost all thinkers and found its apotheosis in modern physics. From the time of the early renaissance to the nineteenth century, physics represented the ultimate expression of scientific investigation for almost all thinkers. Its static laws appeared to be the unchanging principles of all motion and life on earth. By the nineteenth century, it had appeared that only a few details had to be “cleared up” before all science was basically known.

In many ways, this situation changed dramatically with the arrival of Darwinism. It would change even more dramatically in early twentieth-century physics as well. Darwin’s theories of evolution challenged many aspects of the “static” worldview. Even those who did not believe that a divine being created an unchanging world were shaken by the new vistas opened up to science by his studies. It had been a long-accepted inheritance of Western culture to believe that the species of living organisms were unchanging in nature. Though there might be many different kinds of creatures, the kinds themselves were not believed to change. The thesis of a universal morphing of types shattered this cosmology, replacing the old world-view with a totally new one. Among the things that had to change in light of Darwin’s work was the very view of science held by most people.

Consider the underlined sentence. What was the new “cosmology” that arose after Darwin’s day?

Adapted from “Humming-Birds: As Illustrating the Luxuriance of Tropical Nature” in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The food of hummingbirds has been a matter of much controversy. All the early writers down to Buffon believed that they lived solely on the nectar of flowers, but since that time, every close observer of their habits maintains that they feed largely, and in some cases wholly, on insects. Azara observed them on the La Plata in winter taking insects out of the webs of spiders at a time and place where there were no flowers. Bullock, in Mexico, declares that he saw them catch small butterflies, and that he found many kinds of insects in their stomachs. Waterton made a similar statement. Hundreds and perhaps thousands of specimens have since been dissected by collecting naturalists, and in almost every instance their stomachs have been found full of insects, sometimes, but not generally, mixed with a proportion of honey. Many of them in fact may be seen catching gnats and other small insects just like fly-catchers, sitting on a dead twig over water, darting off for a time in the air, and then returning to the twig. Others come out just at dusk, and remain on the wing, now stationary, now darting about with the greatest rapidity, imitating in a limited space the evolutions of the goatsuckers, and evidently for the same end and purpose. Mr. Gosse also remarks, ” All the hummingbirds have more or less the habit, when in flight, of pausing in the air and throwing the body and tail into rapid and odd contortions. This is most observable in the Polytmus, from the effect that such motions have on the long feathers of the tail. That the object of these quick turns is the capture of insects, I am sure, having watched one thus engaged pretty close to me.”

What evidence does Mr. Gosse have to support the claim that hummingbirds eat insects?

Adapted from An Introduction to Astronomy by Forest Ray Moulton (1916 ed.)

It is doubtful if any important scientific idea ever sprang suddenly into the mind of a single man. The great intellectual movements in the world have had long periods of preparation, and often many men were groping for the same truth, without exactly seizing it, before it was fully comprehended.

The foundation on which all science rests is the principle that the universe is orderly, and that all phenomena succeed one another in harmony with invariable laws. Consequently, science was impossible until the truth of this principle was perceived, at least as applied to a limited part of nature.

The phenomena of ordinary observation, as, for example, the weather, depend on such a multitude of factors that it was not easy for men in their primitive state to discover that they occur in harmony with fixed laws. This was the age of superstition, when nature was supposed to be controlled by a great number of capricious gods whose favor could be won by childish ceremonies. Enormous experience was required to dispel such errors and to convince men that the universe is one vast organization whose changes take place in conformity with laws which they can in no way alter.

The actual dawn of science was in prehistoric times, probably in the civilizations that flourished in the valleys of the Nile and the Euphrates. In the very earliest records of these people that have come down to modern times it is found that they were acquainted with many astronomical phenomena and had coherent ideas with respect to the motions of the sun, moon, planets, and stars. It is perfectly clear from their writings that it was from their observations of the heavenly bodies that they first obtained the idea that the universe is not a chaos. Day and night were seen to succeed each other regularly, the moon was found to pass through its phases systematically, the seasons followed one another in order, and in fact the more conspicuous celestial phenomena were observed to occur in an orderly sequence. It is to the glory of astronomy that it first led men to the conclusion that law reigns in the universe.

What does this passage imply to be the effect of not understanding the causes of events?