Adapted from “Darwin’s Predecessors” by J. Arthur Thomson in Evolution in Modern Thought (1917 ed.)

In seeking to discover Darwin's relation to his predecessors, it is useful to distinguish the various services which he rendered to the theory of organic evolution.

As everyone knows, the general idea of the doctrine of descent is that the plants and animals of the present day are the lineal descendants of ancestors on the whole somewhat simpler, that these again are descended from yet simpler forms, and so on backwards towards the literal "Protozoa" and "Protophyta" about which we unfortunately know nothing. Now no one supposes that Darwin originated this idea, which in rudiment at least is as old as Aristotle. What Darwin did was to make it current intellectual coin. He gave it a form that commended itself to the scientific and public intelligence of the day, and he won widespread conviction by showing with consummate skill that it was an effective formula to work with, a key which no lock refused. In a scholarly, critical, and preeminently fair-minded way, admitting difficulties and removing them, foreseeing objections and forestalling them, he showed that the doctrine of descent supplied a modal interpretation of how our present-day fauna and flora have come to be.

In the second place, Darwin applied the evolution-idea to particular problems, such as the descent of man, and showed what a powerful tool it is, introducing order into masses of uncorrelated facts, interpreting enigmas both of structure and function, both bodily and mental, and, best of all, stimulating and guiding further investigation. But here again it cannot be claimed that Darwin was original. The problem of the descent or ascent of man, and other particular cases of evolution, had attracted not a few naturalists before Darwin's day, though no one [except Herbert Spencer in the psychological domain (1855)] had come near him in precision and thoroughness of inquiry.

In the third place, Darwin contributed largely to a knowledge of the factors in the evolution-process, especially by his analysis of what occurs in the case of domestic animals and cultivated plants, and by his elaboration of the theory of natural selection, which Alfred Russel Wallace independently stated at the same time, and of which there had been a few previous suggestions of a more or less vague description. It was here that Darwin's originality was greatest, for he revealed to naturalists the many different forms—often very subtle—which natural selection takes, and with the insight of a disciplined scientific imagination he realized what a mighty engine of progress it has been and is.

What is the overall purpose of this passage?

"Interpreting the Copernican Revolution" by Matthew Minerd (2014)

The expressions of one discipline can often alter the way that other subjects understand themselves. Among such cases are numbered the investigations of Nicolaus Copernicus. Copernicus is best known for his views concerning heliocentrism, a view which eventually obliterated many aspects of the ancient/medieval worldview, at least from the standpoint of physical science. It had always been the natural view of mankind that the earth stood at the center of the universe, a fixed point in reference to the rest of the visible bodies. The sun, stars, and planets all rotated around the earth.

With time, this viewpoint became one of the major reference points for modern life. It provided a provocative image that was used—and often abused—by many people for various purposes. For those who wished to weaken the control of religion on mankind, it was said that the heliocentric outlook proved man’s insignificance. In contrast with earlier geocentrism, heliocentrism was said to show that man is not the center of the universe. He is merely one small being in the midst of a large cosmos. However, others wished to use the “Copernican Revolution” in a very different manner. These thinkers wanted to show that there was another “recentering” that had to happen. Once upon a time, we talked about the world. Now, however, it was necessary to talk of man as the central reference point. Just as the solar system was “centered” on the sun, so too should the sciences be centered on the human person.

However, both of these approaches are fraught with problems. Those who wished to undermine the religious mindset rather misunderstood the former outlook on the solar system. The earlier geocentric mindset did not believe that the earth was the most important body in the heavens. Instead, many ancient and medieval thinkers believed that the highest “sphere” above the earth was the most important being in the physical universe. Likewise, the so-called “Copernican Revolution” in physics was different from the one applied to the human person. Copernicus’ revolution showed that the human point of view was not the center, whereas the later forms of “Copernican revolution” wished to show just the opposite.

Of course, there are many complexities in the history of such important changes in scientific outlook. Nevertheless, it is fascinating to see the wide-reaching effects of such discoveries, even when they have numerous, ambiguous effects.

What can we say, in general, about the interpretations that have been offered for the effects of Copernicus’ discoveries?

"Interpreting the Copernican Revolution" by Matthew Minerd (2014)

The expressions of one discipline can often alter the way that other subjects understand themselves. Among such cases are numbered the investigations of Nicolaus Copernicus. Copernicus is best known for his views concerning heliocentrism, a view which eventually obliterated many aspects of the ancient/medieval worldview, at least from the standpoint of physical science. It had always been the natural view of mankind that the earth stood at the center of the universe, a fixed point in reference to the rest of the visible bodies. The sun, stars, and planets all rotated around the earth.

With time, this viewpoint became one of the major reference points for modern life. It provided a provocative image that was used—and often abused—by many people for various purposes. For those who wished to weaken the control of religion on mankind, it was said that the heliocentric outlook proved man’s insignificance. In contrast with earlier geocentrism, heliocentrism was said to show that man is not the center of the universe. He is merely one small being in the midst of a large cosmos. However, others wished to use the “Copernican Revolution” in a very different manner. These thinkers wanted to show that there was another “recentering” that had to happen. Once upon a time, we talked about the world. Now, however, it was necessary to talk of man as the central reference point. Just as the solar system was “centered” on the sun, so too should the sciences be centered on the human person.

However, both of these approaches are fraught with problems. Those who wished to undermine the religious mindset rather misunderstood the former outlook on the solar system. The earlier geocentric mindset did not believe that the earth was the most important body in the heavens. Instead, many ancient and medieval thinkers believed that the highest “sphere” above the earth was the most important being in the physical universe. Likewise, the so-called “Copernican Revolution” in physics was different from the one applied to the human person. Copernicus’ revolution showed that the human point of view was not the center, whereas the later forms of “Copernican revolution” wished to show just the opposite.

Of course, there are many complexities in the history of such important changes in scientific outlook. Nevertheless, it is fascinating to see the wide-reaching effects of such discoveries, even when they have numerous, ambiguous effects.

Why is the "Copernican revolution" of the human sciences contrary to the literal sense of Copernicus' findings?

"The Place of Lesion Studies in Neuroscience" by Samantha Winter (2013)

It’s easy to forget that the study of neuroscience originated from non-normalized, non-statistically appraised methods like lesion studies. It’s equally easy, with the advent of sophisticated technology, to render such a method obsolete. A small group of neuroscientists today make a case for the reinstitution of lesion studies—the study of abnormal brains with damaged regions in order to better understand the brain—into the twenty-first-century cognitive neuroscience realm. Their suggestion is bold, but their argument is justified.

Cognitive neuroscientists advocate for the use of convergent methods. Many of them argue that with the limitations of our existing techniques, convergent evidence is imperative for sound research. If this is the case, why ignore a method that has potential for implying causality in a domain dominated by correlational research? Rather than advocating for a single method, neuroscientists should take their own advice and use convergent techniques. Sound research should combine a variety of techniques to examine both causal relationships and overcome the individual shortcomings of each method through the use of many.

Lesion studies are also significantly more beneficial now than they were in earlier times. Neuroimaging methods have enhanced our understanding of what contributes to the brain problems most often encountered, and more refined experiments have been developed to confirm the findings from the more unreliable lesion studies. This transformation allows lesion studies to be included alongside the other systems as a mechanism for understanding the human brain.

The primary goal of this passage is __________.

Adapted from Volume Four of The Natural History of Animals: The Animal Life of the World in Its Various Aspects and Relations by James Richard Ainsworth Davis (1903)

The examples of protective resemblance so far quoted are mostly permanent adaptations to one particular sort of surrounding. There are, however, numerous animals which possess the power of adjusting their color more or less rapidly so as to harmonize with a changing environment.

Some of the best known of these cases are found among those mammals and birds that inhabit countries more or less covered with snow during a part of the year. A good instance is afforded by the Irish or variable hare, which is chiefly found in Ireland and Scotland. In summer, this looks very much like an ordinary hare, though rather grayer in tint and smaller in size, but in winter it becomes white with the exception of the black tips to the ears. Investigations that have been made on the closely allied American hare seem to show that the phenomenon is due to the growth of new hairs of white hue. 

The common stoat is subject to similar color change in the northern parts of its range. In summer it is of a bright reddish brown color with the exception of the under parts, which are yellowish white, and the end of the tail, which is black. But in winter, the entire coat, save only the tip of the tail, becomes white, and in that condition the animal is known as an ermine. A similar example is afforded by the weasel. The seasonal change in the vegetarian Irish hare is purely of protective character, but in such an actively carnivorous creature as a stoat or weasel, it is aggressive as well, rendering the animal inconspicuous to its prey.

Which of the following best expresses the main idea of the passage?

Adapted from Volume Four of The Natural History of Animals: The Animal Life of the World in Its Various Aspects and Relations by James Richard Ainsworth Davis (1903)

The examples of protective resemblance so far quoted are mostly permanent adaptations to one particular sort of surrounding. There are, however, numerous animals which possess the power of adjusting their color more or less rapidly so as to harmonize with a changing environment.

Some of the best known of these cases are found among those mammals and birds that inhabit countries more or less covered with snow during a part of the year. A good instance is afforded by the Irish or variable hare, which is chiefly found in Ireland and Scotland. In summer, this looks very much like an ordinary hare, though rather grayer in tint and smaller in size, but in winter it becomes white with the exception of the black tips to the ears. Investigations that have been made on the closely allied American hare seem to show that the phenomenon is due to the growth of new hairs of white hue. 

The common stoat is subject to similar color change in the northern parts of its range. In summer it is of a bright reddish brown color with the exception of the under parts, which are yellowish white, and the end of the tail, which is black. But in winter, the entire coat, save only the tip of the tail, becomes white, and in that condition the animal is known as an ermine. A similar example is afforded by the weasel. The seasonal change in the vegetarian Irish hare is purely of protective character, but in such an actively carnivorous creature as a stoat or weasel, it is aggressive as well, rendering the animal inconspicuous to its prey.

The purpose of the passage’s third paragraph is __________.

Adapted from “Feathers of Sea Birds and Wild Fowl for Bedding” from The Utility of Birds by Edward Forbush (ed. 1922)

In the colder countries of the world, the feathers and down of waterfowl have been in great demand for centuries as filling for beds and pillows. Such feathers are perfect non-conductors of heat, and beds, pillows, or coverlets filled with them represent the acme of comfort and durability. The early settlers of New England saved for such purposes the feathers and down from the thousands of wild-fowl which they killed, but as the population increased in numbers, the quantity thus furnished was insufficient, and the people sought a larger supply in the vast colonies of ducks and geese along the Labrador coast. 

The manner in which the feathers and down were obtained, unlike the method practiced in Iceland, did not tend to conserve and protect the source of supply. In Iceland, the people have continued to receive for many years a considerable income by collecting eider down, but there they do not “kill the goose that lays the golden eggs.” Ducks line their nests with down plucked from their own breasts and that of the eider is particularly valuable for bedding. In Iceland, these birds are so carefully protected that they have become as tame and unsuspicious as domestic fowls In North America. Where they are constantly hunted they often conceal their nests in the midst of weeds or bushes, but in Iceland, they make their nests and deposit their eggs in holes dug for them in the sod. A supply of the ducks is maintained so that the people derive from them an annual income.

In North America, quite a different policy was pursued. The demand for feathers became so great in the New England colonies about the middle of the eighteenth century that vessels were fitted out there for the coast of Labrador for the express purpose of securing the feathers and down of wild fowl. Eider down having become valuable and these ducks being in the habit of congregating by thousands on barren islands of the Labrador coast, the birds became the victims of the ships’ crews. As the ducks molt all their primary feathers at once in July or August and are then quite incapable of flight and the young birds are unable to fly until well grown, the hunters were able to surround the helpless birds, drive them together, and kill them with clubs. Otis says that millions of wildfowl were thus destroyed and that in a few years their haunts were so broken up by this wholesale slaughter and their numbers were so diminished that feather voyages became unprofitable and were given up. 

This practice, followed by the almost continual egging, clubbing, shooting, etc. by Labrador fishermen, may have been a chief factor in the extinction of the Labrador duck, that species of supposed restricted breeding range. No doubt had the eider duck been restricted in its breeding range to the islands of Labrador, it also would have been exterminated long ago.

Which of the following best states the main idea of this passage?

Adapted from "Recent Views as to Direct Action of Light on the Colors of Flowers and Fruits" in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The theory that the brilliant colors of flowers and fruits is due to the direct action of light has been supported by a recent writer by examples taken from the arctic instead of from the tropical flora. In the arctic regions, vegetation is excessively rapid during the short summer, and this is held to be due to the continuous action of light throughout the long summer days. “The further we advance towards the north, the more the leaves of plants increase in size as if to absorb a greater proportion of the solar rays. M. Grisebach says that during a journey in Norway he observed that the majority of deciduous trees had already, at the 60th degree of latitude, larger leaves than in Germany, while M. Ch. Martins has made a similar observation as regards the leguminous plants cultivated in Lapland.” The same writer goes on to say that all the seeds of cultivated plants acquire a deeper color the further north they are grown, white haricots becoming brown or black, and white wheat becoming brown, while the green color of all vegetation becomes more intense. The flowers also are similarly changed: those which are white or yellow in central Europe becoming red or orange in Norway. This is what occurs in the Alpine flora, and the cause is said to be the same in both—the greater intensity of the sunlight. In the one the light is more persistent, in the other more intense because it traverses a less thickness of atmosphere.

Admitting the facts as above stated to be in themselves correct, they do not by any means establish the theory founded on them; and it is curious that Grisebach, who has been quoted by this writer for the fact of the increased size of the foliage, gives a totally different explanation of the more vivid colors of Arctic flowers. He says, “We see flowers become larger and more richly colored in proportion as, by the increasing length of winter, insects become rarer, and their cooperation in the act of fecundation is exposed to more uncertain chances.” (Vegetation du Globe, col. i. p. 61—French translation.) This is the theory here adopted to explain the colors of Alpine plants, and we believe there are many facts that will show it to be the preferable one. The statement that the white and yellow flowers of temperate Europe become red or golden in the Arctic regions must we think be incorrect. By roughly tabulating the colors of the plants given by Sir Joseph Hooker as permanently Arctic, we find among fifty species with more or less conspicuous flowers, twenty-five white, twelve yellow, eight purple or blue, three lilac, and two red or pink; showing a very similar proportion of white and yellow flowers to what obtains further south.

The purpose of this passage is ___________.

Adapted from "Recent Views as to Direct Action of Light on the Colors of Flowers and Fruits" in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The theory that the brilliant colors of flowers and fruits is due to the direct action of light has been supported by a recent writer by examples taken from the arctic instead of from the tropical flora. In the arctic regions, vegetation is excessively rapid during the short summer, and this is held to be due to the continuous action of light throughout the long summer days. "The further we advance towards the north, the more the leaves of plants increase in size as if to absorb a greater proportion of the solar rays. M. Grisebach says that during a journey in Norway he observed that the majority of deciduous trees had already, at the 60th degree of latitude, larger leaves than in Germany, while M. Ch. Martins has made a similar observation as regards the leguminous plants cultivated in Lapland.” The same writer goes on to say that all the seeds of cultivated plants acquire a deeper color the further north they are grown, white haricots becoming brown or black, and white wheat becoming brown, while the green color of all vegetation becomes more intense. The flowers also are similarly changed: those which are white or yellow in central Europe becoming red or orange in Norway. This is what occurs in the Alpine flora, and the cause is said to be the same in both—the greater intensity of the sunlight. In the one the light is more persistent, in the other more intense because it traverses a less thickness of atmosphere.

Admitting the facts as above stated to be in themselves correct, they do not by any means establish the theory founded on them; and it is curious that Grisebach, who has been quoted by this writer for the fact of the increased size of the foliage, gives a totally different explanation of the more vivid colors of Arctic flowers. He says, “We see flowers become larger and more richly colored in proportion as, by the increasing length of winter, insects become rarer, and their cooperation in the act of fecundation is exposed to more uncertain chances.” (Vegetation du Globe, col. i. p. 61—French translation.) This is the theory here adopted to explain the colors of Alpine plants, and we believe there are many facts that will show it to be the preferable one. The statement that the white and yellow flowers of temperate Europe become red or golden in the Arctic regions must we think be incorrect. By roughly tabulating the colors of the plants given by Sir Joseph Hooker as permanently Arctic, we find among fifty species with more or less conspicuous flowers, twenty-five white, twelve yellow, eight purple or blue, three lilac, and two red or pink; showing a very similar proportion of white and yellow flowers to what obtains further south.

The author’s critique of the theory presented in the first paragraph is that __________.

Adapted from "Taking a Second Look: An Analysis of Genetic Markers in Species Relatedness" by Joseph Ritchie (2014)

Phylogenetics is the study of genetic composition in various species and is used by evolutionary biologists to investigate similarities in the molecular sequences of proteins in varying organisms. The amino acid sequences that build proteins are used to construct mathmatical matrices that aid in determining evolutionary ties through the investigation of percentage similarities. The study of these matrices helps to expose evolutionary relationships between species that may not have the same overt characteristics.

Species adapt and evolve based on the pressures that exist in their environment. Climate, food source, and habitat availability are only a few factors that act on species adaptation. These stressors can alter the physical characteristics of organisms. This divergence in evolution has made it difficult to determine the interrelatedness of organisms by analyzing their physical characteristics alone.

For instance, looking only at physical characteristics, the ghost bat resembles a pigeon more than a spider monkey; however, phylogenetics has found that the amino acid sequences that construct the beta hemoglobin molecules of bats are twenty percent more similar to those of mammalian primates than those of birds. This helps reject the assumption that common physical characteristics between species are all that is needed to determine relatedness. 

The differences produced by divergent evolution observed in the forest-dwelling, arboreal spider monkey and the nocturnal, airborne ghost bat can be reconciled through homology. Homologous characteristics are anatomical traits that are similar in two or more different species. For instance, the bone structure of a spider monkey’s wrist and fingers greatly resembles that of a bat’s wing or even a whale’s fin. These similarities are reinforced by phylogenetic evidence that supports the idea that physically dissimilar species can be evolutionarily related through anatomical and genetic similarities.

The central idea of this passage is best described by which of the following statements?