All SAT Critical Reading Resources
Example Questions
Example Question #3 : Science Passages
"The Multiple Sides of Computer Science" by Matthew Minerd (2014)
It often takes some time for a new discipline to become recognized as an independent science. An excellent example of this is computer science. In many ways, this science still is a hodgepodge of several different sciences, each one having its own distinct character. For example, some computer scientists are almost indistinguishable from mathematicians. Many of the most difficult topics in pattern recognition and data communications require intensive mathematics in order to provide software solutions. Years of training in the appropriate disciplines are necessary before the computer scientist can even begin to work as a programmer in such areas. In contrast to those computer scientists who work with complex mathematics, many computer scientists work on areas of hardware development that are similar to disciplines like electrical engineering and physics.
However, computer science has its own particular problems regarding the unity of its subject matter. There are many practical applications for computing work; therefore, many computer scientists focus on learning a large set of skills in programming languages, development environments, and even information technology. All of these disciplines have a certain practical coloration that is quite distinct from the theoretical concepts used in other parts of the field. Nevertheless, these practical topics add to the broad range of topics covered by most academic programs that claim to focus on “computer science.” It can only be hoped that these disciplines will increase in orderliness in the coming decades.
What is the main point introduced in the second paragraph?
None of these
Computer science only becomes "real" when it is applied to the real world
Many programmers are mere technicians, not real computer scientists
Computer science is a very practical science for most topics
The applied nature of computer science creates a unique set of problems regarding the discipline's unity
The applied nature of computer science creates a unique set of problems regarding the discipline's unity
The second paragraph focuses on the fact that computer science has a host of practical applications. In particular, these practical applications make it even more difficult to see the focus of computer science studies. Therefore, they tend to hide the unity of the topic even more (that is, even beyond the more "academic" issues discussed in the first paragraph).
Example Question #1 : Ideas In Science Passages
"Abstraction in the Sciences" by Matthew Minerd (2014)
Thinking “abstractly” is not a term that means quite the same thing in all of the sciences. Although we rarely think about this, it plays a key role in almost all of our day-to-day thought. Consider a zoologist working in a lab with many animals. When she is studying any individual tiger, she is not completely worried about the particular tiger—at least not primarily. Instead, she is trying to figure out certain characteristics of tigers in general. By meticulous testing, the zoologist carefully works out the physiology of tigers and considers what are absolutely necessary elements of their physical makeup. Even when she places a tiger in different habitats, her sight is aimed at the general condition of tigers and their needs in general.
However, things become even stranger when you start to consider how we think about mathematical objects. Consider the case of geometric figures. A triangle appears to be rather simple for most of us to think about. You can draw a triangle on a piece of paper, each side having a certain thickness and length. However when you think about this in geometry class, the triangle’s edges have no real thickness. Neither a point nor a line has a thickness for the mathematician. Such a thickness only exists on our paper, which represents the point or line. Consider also a line drawn on a piece of graph paper. Technically, there are an infinite number of points in the line. Indeed, even between 4.5 and 4.6, there are an infinite number of numbers—for example 4.55 is between them, then 4.555 between 4.55 and 4.6, and 4.5555 between 4.555 and 4.6, et cetera. In all of these cases, the mathematical reality takes on a very peculiar character when you consider it in the abstract. However, the concrete triangle remains very tangible and ordinary. Likewise, 4.6 and 4.5 inches still have 0.1 inches between them. Nevertheless, in the abstract, mathematical realities are quite strange, even stranger then the idea of “a tiger in general.”
Which of the following would strengthen the ending of the first paragraph?
"Of course, she becomes quite attached to all of the animals in her care."
"Every animal is carefully tested as a potential test subject."
None of the other answers
"Each individual tiger has its own unique characteristics and abilities."
"The details of any particular tiger’s life are interesting only as a means to this end."
"The details of any particular tiger’s life are interesting only as a means to this end."
As written, the last sentence in the first paragraph states that the scientist is interested in figuring out what "tigers in general" are like. Therefore, any particular tiger is not as important as this general nature of tigers (how they generally can live, thrive, etc). The best way to end this paragraph is by reiterating this point, which is what the correct answer does.
Example Question #2 : Science Passages
Adapted from An Introduction to Astronomy by Forest Ray Moulton (1916 ed.)
The ancient Greeks, at a period four or five hundred years preceding the common era, definitely undertook to find from systematic observation how celestial phenomena follow one another. They determined very accurately the number of days in the year, the period of the moon's revolution, and the paths of the sun and the moon among the stars; they correctly explained the cause of eclipses and learned how to predict them with a considerable degree of accuracy; they undertook to measure the distances to the heavenly bodies, and to work out a complete system that would represent their motions. The idea was current among the Greek philosophers that the earth was spherical, that it turned on its axis, and, among some of them, that it revolved around the sun. They had true science in the modern acceptance of the term, but it was largely confined to the relations among celestial phenomena.
The conception that the heavens are orderly, which they definitely formulated and acted on with remarkable success, has been extended, especially in the last two centuries, so as to include the whole universe. The extension was first made to the inanimate world and then to the more complicated phenomena associated with living beings. Every increase in carefully recorded experience has confirmed and strengthened the belief that nature is perfectly orderly, until now every one who has had an opportunity of becoming familiar with any science is firmly convinced of the truth of this principle, which is the basis of all science.
What is the main idea of the second paragraph?
That science has finally outlined the complete details of the orderliness of the universe
That science was only possible because of the orderliness of the heavens
None of the other answers
That science depends upon the regularity of that which it studies
That the Greeks could not fathom the depths of the universe's orderliness
That science depends upon the regularity of that which it studies
The general idea of the second paragraph is that orderliness is necessary for there to be any science, for science discovers laws and enumerates them. The paragraph goes on to stress the growth in this awareness. However, it does not say that every detail has been so recognized. Likewise, it does not speak disparagingly of the Greeks, though they did not realize the extent to which modern physics would assert order in the whole of the cosmos. The most direct answer is the best: "That science depends upon the regularity of that which it studies."
Example Question #11 : Analyzing Main Idea, Theme, And Purpose In Natural Science Passages
Adapted from An Introduction to Astronomy by Forest Ray Moulton (1916 ed.)
The ancient Greeks, at a period four or five hundred years preceding the common era, definitely undertook to find from systematic observation how celestial phenomena follow one another. They determined very accurately the number of days in the year, the period of the moon's revolution, and the paths of the sun and the moon among the stars; they correctly explained the cause of eclipses and learned how to predict them with a considerable degree of accuracy; they undertook to measure the distances to the heavenly bodies, and to work out a complete system that would represent their motions. The idea was current among the Greek philosophers that the earth was spherical, that it turned on its axis, and, among some of them, that it revolved around the sun. They had true science in the modern acceptance of the term, but it was largely confined to the relations among celestial phenomena.
The conception that the heavens are orderly, which they definitely formulated and acted on with remarkable success, has been extended, especially in the last two centuries, so as to include the whole universe. The extension was first made to the inanimate world and then to the more complicated phenomena associated with living beings. Every increase in carefully recorded experience has confirmed and strengthened the belief that nature is perfectly orderly, until now every one who has had an opportunity of becoming familiar with any science is firmly convinced of the truth of this principle, which is the basis of all science.
What is the purpose of the underlined sentence, "The extension was first made . . ."?
To contrast developments in Greek and modern science
None of the other answers
To show the many ways that science could extend Greek concepts
To lay out a plan for the extension of science into new domains
To describe the history of late Greek science
None of the other answers
This sentence helps to show how the initial optimism of Greek "astronomy" was able to be extended to many other topics, noting the regularity found in many other phenomena. At best, this could be considered a history of modern science's development, though it would be a rather thin history with few real details. None of the answers really state this. The closest might be, "To contrast developments in Greek and modern science." However, the sentence really is not contrasting the two.
Example Question #12 : Analyzing Main Idea, Theme, And Purpose In Natural Science Passages
Adapted from “Humming-Birds: As Illustrating the Luxuriance of Tropical Nature” in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)
The food of hummingbirds has been a matter of much controversy. All the early writers down to Buffon believed that they lived solely on the nectar of flowers, but since that time, every close observer of their habits maintains that they feed largely, and in some cases wholly, on insects. Azara observed them on the La Plata in winter taking insects out of the webs of spiders at a time and place where there were no flowers. Bullock, in Mexico, declares that he saw them catch small butterflies, and that he found many kinds of insects in their stomachs. Waterton made a similar statement. Hundreds and perhaps thousands of specimens have since been dissected by collecting naturalists, and in almost every instance their stomachs have been found full of insects, sometimes, but not generally, mixed with a proportion of honey. Many of them in fact may be seen catching gnats and other small insects just like fly-catchers, sitting on a dead twig over water, darting off for a time in the air, and then returning to the twig. Others come out just at dusk, and remain on the wing, now stationary, now darting about with the greatest rapidity, imitating in a limited space the evolutions of the goatsuckers, and evidently for the same end and purpose. Mr. Gosse also remarks, ” All the hummingbirds have more or less the habit, when in flight, of pausing in the air and throwing the body and tail into rapid and odd contortions. This is most observable in the Polytmus, from the effect that such motions have on the long feathers of the tail. That the object of these quick turns is the capture of insects, I am sure, having watched one thus engaged pretty close to me.”
The purpose of this passage is __________.
to propose a definitive experiment about what hummingbirds eat
to discuss the Polytmus’ feeding habits
to explain why one should feed a captive hummingbird insects and not flower nectar
to consider the opinions of scientists on what hummingbirds eat
to critique the opinions of other scientists
to consider the opinions of scientists on what hummingbirds eat
When answering questions about a passage’s purpose or main idea, it’s important to pick an answer choice that is broad enough to encompass the entire passage. For instance, while the Polytmus’ feeding habits are discussed in the passage, it can’t be said to be the purpose of the passage, since it’s only mentioned as a small detail. The passage can’t be said to critique the opinions of other scientists, because for the most part, the author quotes findings by scientists with whom he does not disagree. A definitive experiment is never proposed, and captive hummingbirds are never discussed. The passage does provide the opinions of scientists on what hummingbirds eat; this accurately captures the intent of the entire passage, not just some of its parts, and doesn’t state it too broadly.
Example Question #13 : Natural Sciences
Adapted from “Darwin’s Predecessors” by J. Arthur Thomson in Evolution in Modern Thought (1917 ed.)
In seeking to discover Darwin's relation to his predecessors, it is useful to distinguish the various services which he rendered to the theory of organic evolution.
As everyone knows, the general idea of the doctrine of descent is that the plants and animals of the present day are the lineal descendants of ancestors on the whole somewhat simpler, that these again are descended from yet simpler forms, and so on backwards towards the literal "Protozoa" and "Protophyta" about which we unfortunately know nothing. Now no one supposes that Darwin originated this idea, which in rudiment at least is as old as Aristotle. What Darwin did was to make it current intellectual coin. He gave it a form that commended itself to the scientific and public intelligence of the day, and he won widespread conviction by showing with consummate skill that it was an effective formula to work with, a key which no lock refused. In a scholarly, critical, and preeminently fair-minded way, admitting difficulties and removing them, foreseeing objections and forestalling them, he showed that the doctrine of descent supplied a modal interpretation of how our present-day fauna and flora have come to be.
In the second place, Darwin applied the evolution-idea to particular problems, such as the descent of man, and showed what a powerful tool it is, introducing order into masses of uncorrelated facts, interpreting enigmas both of structure and function, both bodily and mental, and, best of all, stimulating and guiding further investigation. But here again it cannot be claimed that Darwin was original. The problem of the descent or ascent of man, and other particular cases of evolution, had attracted not a few naturalists before Darwin's day, though no one [except Herbert Spencer in the psychological domain (1855)] had come near him in precision and thoroughness of inquiry.
In the third place, Darwin contributed largely to a knowledge of the factors in the evolution-process, especially by his analysis of what occurs in the case of domestic animals and cultivated plants, and by his elaboration of the theory of natural selection, which Alfred Russel Wallace independently stated at the same time, and of which there had been a few previous suggestions of a more or less vague description. It was here that Darwin's originality was greatest, for he revealed to naturalists the many different forms—often very subtle—which natural selection takes, and with the insight of a disciplined scientific imagination he realized what a mighty engine of progress it has been and is.
What is the overall purpose of this passage?
To list some theories with which Darwin disagreed
None of the other answers
To defend Darwin's account of evolution against those who deny its cogency
To praise every aspect of Darwin's account of evolution
To provide comprehensive account of the history of Darwin's thought
None of the other answers
In this selection, the main idea is stated in the very first sentence, so long as you pay attention to it: "In seeking to discover Darwin's relation to his predecessors, it is useful to distinguish the various services which he rendered to the theory of organic evolution." The passage is directing our attention not to his predecessors per se. It is presenting "the various services which he rendered to the theory of organic evolution." It lists several such "services" that Darwin rendered in order to show how he helped to develop the theory of evolution in biology. None of the answers state this adequately.
Example Question #13 : Analyzing Main Idea, Theme, And Purpose In Natural Science Passages
"Cacti" by Ami Dave (2013)
Cacti are plants suited to the desert, and we must always keep this factor in mind when growing ornamental cacti in our gardens, for it helps us provide cacti with conditions that allow them to survive and thrive. For example, a cactus should never be watered over its body, as it will start to rot. This is because it is covered with a waxy coating which prevents water loss through evaporation. When one waters the cactus over its body, the waxy coating is washed away and the plant begins to rot. The amount of water that one must supply to the cactus is very much dependent upon the season and upon the climate of the place. During the summer season one should water cacti every four days, whereas in the rainy season, once every fifteen days is quite enough.
Cacti need a minimum of two and a half hours of sunlight per day; however, they should not be kept in the sun all day because they may wrinkle when exposed to too much bright sunlight. Unlike other plants, cacti produce carbon dioxide during the day and oxygen during the night, so they are ideal plants to be kept in bedrooms to freshen up the air at night.
If a cactus is to thrive and prosper, the size of the pot in which it is grown needs to be monitored carefully. The pot should always be a little smaller than the plant itself because it is only when the plant has to struggle to survive that it will thrive. If the pot is too spacious and the plant does not need to struggle, chances are that the cactus will die. Similarly, if a cactus shows no signs of growth, stop watering it. Watering should be resumed only when the plant begins to grow again.
The substrata of a cactus pot is ideally composed of pieces of broken bricks at the bottom, followed by a layer of charcoal above the bricks, and then coarse sand and pebbles above the charcoal. Leaf mould is the best manure.
Grafting cacti is very simple. A very small piece of the cactus plant should be stuck with tape to the plant that needs grafting. The smaller the piece, the easier it is to graft. To reproduce cacti, one has to simply cut off a piece of the cactus, allow it to dry for a few days, and then place it over the cacti substrate. It will automatically develop roots.
It is very easy to differentiate between cacti and other plants that look like cacti. All cacti have fine hair at the base of each thorn. The so-called “thorns” are in fact highly modified leaves which prevent loss of water through transpiration. If one ever gets pricked by cacti thorns, one should take tape, place it over the area where the thorns have penetrated the skin, and then peel it off. All of the thorns will get stuck to the tape and will be removed.
The purpose of this passage is to __________.
outline the differences between cacti and other plants
explain how to correctly graft cacti
explain the proper conditions and protocols for growing cacti
describe the physical characteristics of a cactus
explain what to do if you prick yourself on cactus thorns
explain the proper conditions and protocols for growing cacti
Cacti grafts, physical characteristics of cacti, what to do if you prick yourself on cactus thorns, and differences between cacti and other plants are all ideas that are discussed in the passage; however, they are not the MAIN purpose of the passage. They are all details that comprise the paragraphs. The purpose of the passage is to explain the proper conditions and protocols for growing cacti.
Example Question #21 : Critical Comprehension
Adapted from “Birds in Retreat” in “Animal Defences—Active Defence” in Volume Four of The Natural History of Animals: The Animal Life of the World in Its Various Aspects and Relations by James Richard Ainsworth Davis (1903)
Among the large running birds are forms, like the African ostrich, in which the absence of powers of flight is largely compensated by the specialization of the legs for the purpose of rapid movement on the ground. For straightforward retreat in open country nothing could be more effective; but another kind of adaptation is required in birds like rails, which are deficient in powers of flight, and yet are able to run through thickly-growing vegetation with such rapidity as to commonly elude their enemies. This is rendered possible by the shape of their bodies, which are relatively narrow and flattened from side to side, so as to easily slip between the stems of grasses, rushes, and similar plants. Anyone who has pursued our native land-rail or corn-crake with intent to capture will have noted how extremely difficult it is even to get within sight of a bird of this sort.
Certain birds, unfortunately for themselves, have lost the power of flight without correspondingly increased powers of running, and have paid the penalty of extinction. Such an arrangement, as might be anticipated, was the result of evolution in islands devoid of any predatory ground-animals, and a classic example of it is afforded by the dodo and its allies, birds related to the pigeons. The dodo itself was a large and clumsy-looking species that at one time abounded in the island of Mauritius, which, like oceanic islands generally, possessed no native mammals, while its indigenous reptiles were only represented by lizards. The ubiquitous sailor, however, and the animals (especially swine) which he introduced, brought about the extinction of this helpless bird in less than a century after its first discovery in 1598. Its memory is now only kept green by a few contemporary drawings and descriptions, certain museum remains, and the proverb "as extinct as a dodo.” A similar fate must overtake any organism suddenly exposed to new and unfavorable conditions, if devoid of sufficient plasticity to rapidly accommodate itself to the altered environment.
Which of the following best states the main idea of this passage?
Birds employ a variety of defenses for avoiding predators.
Flightless birds adapt to their particular surroundings.
Flightless birds sometimes develop strong running abilities, but if they don’t, they may be threatened with extinction.
The extinction of the dodo can be traced to human causes.
The African ostrich’s strong leg muscles makes it well adapted to its environment.
Flightless birds sometimes develop strong running abilities, but if they don’t, they may be threatened with extinction.
When asked to identify a passage’s main idea, it is important to pick out an answer choice to which all of the paragraphs can relate, but that is not too broad in including things that the passage does not discuss. Considering if each of the answer choices falls into one or another of these categories can help you narrow down your choices. For instance, “Birds employ a variety of defenses for avoiding predators” is far too broad to accurately describer this passage’s main idea. The author only discusses flightless birds, not all birds. “Flightless birds adapt to their particular surroundings” cannot be correct either, as the first paragraph discusses this, but the second paragraph discusses a flightless bird that did not adapt to its surroundings. Since the second paragraph can’t relate to this answer, it can’t be the main idea of the entire passage. Two of the remaining answer choices can be discarded due to their being too detailed: “The African ostrich’s strong leg muscles makes it well adapted to its environment” and “The extinction of the dodo can be traced to human causes.” While the first of these choices is stated in the first paragraph and the second is stated in the second paragraph, neither relates to the other paragraph, or even adequately summarizes the entire paragraph in which it appears. This leaves us with one answer choice, the correct one: “Flightless birds sometimes develop strong running abilities, but if they don’t, they may be threatened with extinction.” Each of the two paragraphs can relate to this answer choice, but it doesn’t include things that the passage doesn’t discuss.
Example Question #1 : Natural Sciences
Adapted from The Effects of Cross & Self-Fertilisation in the Vegetable Kingdom by Charles Darwin (1876)
As it is impossible to exclude such minute pollen-carrying insects as Thrips, flowers which it was intended to fertilise with their own pollen may sometimes have been afterwards crossed with pollen brought by these insects from another flower on the same plant; but as we shall hereafter see, a cross of this kind does not produce any effect, or at most only a slight one. When two or more plants were placed near one another under the same net, as was often done, there is some real though not great danger of the flowers which were believed to be self-fertilised being afterwards crossed with pollen brought by Thrips from a distinct plant. I have said that the danger is not great because I have often found that plants which are self-sterile, unless aided by insects, remained sterile when several plants of the same species were placed under the same net. If, however, the flowers which had been presumably self-fertilised by me were in any case afterwards crossed by Thrips with pollen brought from a distinct plant, crossed seedlings would have been included amongst the self-fertilised; but it should be especially observed that this occurrence would tend to diminish and not to increase any superiority in average height, fertility, etc., of the crossed over the self-fertilised plants.
As the flowers which were crossed were never castrated, it is probable or even almost certain that I sometimes failed to cross-fertilise them effectually, and that they were afterwards spontaneously self-fertilised. This would have been most likely to occur with dichogamous species, for without much care it is not easy to perceive whether their stigmas are ready to be fertilised when the anthers open. But in all cases, as the flowers were protected from wind, rain, and the access of insects, any pollen placed by me on the stigmatic surface whilst it was immature, would generally have remained there until the stigma was mature; and the flowers would then have been crossed as was intended. Nevertheless, it is highly probable that self-fertilised seedlings have sometimes by this means got included amongst the crossed seedlings. The effect would be, as in the former case, not to exaggerate but to diminish any average superiority of the crossed over the self-fertilised plants.
Errors arising from the two causes just named, and from others,—such as some of the seeds not having been thoroughly ripened, though care was taken to avoid this error—the sickness or unperceived injury of any of the plants,—will have been to a large extent eliminated, in those cases in which many crossed and self-fertilised plants were measured and an average struck. Some of these causes of error will also have been eliminated by the seeds having been allowed to germinate on bare damp sand, and being planted in pairs; for it is not likely that ill-matured and well-matured, or diseased and healthy seeds, would germinate at exactly the same time. The same result will have been gained in the several cases in which only a few of the tallest, finest, and healthiest plants on each side of the pots were measured.
Kolreuter and Gartner have proved that with some plants several, even as many as from fifty to sixty, pollen-grains are necessary for the fertilisation of all the ovules in the ovarium. Naudin also found in the case of Mirabilis that if only one or two of its very large pollen-grains were placed on the stigma, the plants raised from such seeds were dwarfed. I was therefore careful to give an amply sufficient supply of pollen, and generally covered the stigma with it; but I did not take any special pains to place exactly the same amount on the stigmas of the self-fertilised and crossed flowers. After having acted in this manner during two seasons, I remembered that Gartner thought, though without any direct evidence, that an excess of pollen was perhaps injurious. It was therefore necessary to ascertain whether the fertility of the flowers was affected by applying a rather small and an extremely large quantity of pollen to the stigma. Accordingly a very small mass of pollen-grains was placed on one side of the large stigma in sixty-four flowers of Ipomoea purpurea, and a great mass of pollen over the whole surface of the stigma in sixty-four other flowers. In order to vary the experiment, half the flowers of both lots were on plants produced from self-fertilised seeds, and the other half on plants from crossed seeds. The sixty-four flowers with an excess of pollen yielded sixty-one capsules; and excluding four capsules, each of which contained only a single poor seed, the remainder contained on an average 5.07 seeds per capsule. The sixty-four flowers with only a little pollen placed on one side of the stigma yielded sixty-three capsules, and excluding one from the same cause as before, the remainder contained on an average 5.129 seeds. So that the flowers fertilised with little pollen yielded rather more capsules and seeds than did those fertilised with an excess; but the difference is too slight to be of any significance. On the other hand, the seeds produced by the flowers with an excess of pollen were a little heavier of the two; for 170 of them weighed 79.67 grains, whilst 170 seeds from the flowers with very little pollen weighed 79.20 grains. Both lots of seeds having been placed on damp sand presented no difference in their rate of germination. We may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.
What is the main idea of the passage?
An evaluation of the confounding factors of the experiment.
An assessment of the effect of Thrips on fertilisation of plants.
A discussion of the different ways in which flowers can be pollinated.
A description of the methods used to artificially pollinate different plants.
An argument over the best ways to germinate different plants.
An evaluation of the confounding factors of the experiment.
The confounding factors of the experiment are those which may have had an adverse affect on the result. This passage evaluates different confounding factors and assesses how steps to prevent them were taken or verifies if they would have had a great enough impact on the results for them to be taken into consideration. The third paragraph supports this answer, as it starts “Errors arising from the two causes just named, and from others” suggesting that that is what the author is talking about in the whole passage rather than the specifics of each paragraph.
Example Question #2 : Natural Sciences
Adapted from The Effects of Cross & Self-Fertilisation in the Vegetable Kingdom by Charles Darwin (1876)
As it is impossible to exclude such minute pollen-carrying insects as Thrips, flowers which it was intended to fertilise with their own pollen may sometimes have been afterwards crossed with pollen brought by these insects from another flower on the same plant; but as we shall hereafter see, a cross of this kind does not produce any effect, or at most only a slight one. When two or more plants were placed near one another under the same net, as was often done, there is some real though not great danger of the flowers which were believed to be self-fertilised being afterwards crossed with pollen brought by Thrips from a distinct plant. I have said that the danger is not great because I have often found that plants which are self-sterile, unless aided by insects, remained sterile when several plants of the same species were placed under the same net. If, however, the flowers which had been presumably self-fertilised by me were in any case afterwards crossed by Thrips with pollen brought from a distinct plant, crossed seedlings would have been included amongst the self-fertilised; but it should be especially observed that this occurrence would tend to diminish and not to increase any superiority in average height, fertility, etc., of the crossed over the self-fertilised plants.
As the flowers which were crossed were never castrated, it is probable or even almost certain that I sometimes failed to cross-fertilise them effectually, and that they were afterwards spontaneously self-fertilised. This would have been most likely to occur with dichogamous species, for without much care it is not easy to perceive whether their stigmas are ready to be fertilised when the anthers open. But in all cases, as the flowers were protected from wind, rain, and the access of insects, any pollen placed by me on the stigmatic surface whilst it was immature, would generally have remained there until the stigma was mature; and the flowers would then have been crossed as was intended. Nevertheless, it is highly probable that self-fertilised seedlings have sometimes by this means got included amongst the crossed seedlings. The effect would be, as in the former case, not to exaggerate but to diminish any average superiority of the crossed over the self-fertilised plants.
Errors arising from the two causes just named, and from others,—such as some of the seeds not having been thoroughly ripened, though care was taken to avoid this error—the sickness or unperceived injury of any of the plants,—will have been to a large extent eliminated, in those cases in which many crossed and self-fertilised plants were measured and an average struck. Some of these causes of error will also have been eliminated by the seeds having been allowed to germinate on bare damp sand, and being planted in pairs; for it is not likely that ill-matured and well-matured, or diseased and healthy seeds, would germinate at exactly the same time. The same result will have been gained in the several cases in which only a few of the tallest, finest, and healthiest plants on each side of the pots were measured.
Kolreuter and Gartner have proved that with some plants several, even as many as from fifty to sixty, pollen-grains are necessary for the fertilisation of all the ovules in the ovarium. Naudin also found in the case of Mirabilis that if only one or two of its very large pollen-grains were placed on the stigma, the plants raised from such seeds were dwarfed. I was therefore careful to give an amply sufficient supply of pollen, and generally covered the stigma with it; but I did not take any special pains to place exactly the same amount on the stigmas of the self-fertilised and crossed flowers. After having acted in this manner during two seasons, I remembered that Gartner thought, though without any direct evidence, that an excess of pollen was perhaps injurious. It was therefore necessary to ascertain whether the fertility of the flowers was affected by applying a rather small and an extremely large quantity of pollen to the stigma. Accordingly a very small mass of pollen-grains was placed on one side of the large stigma in sixty-four flowers of Ipomoea purpurea, and a great mass of pollen over the whole surface of the stigma in sixty-four other flowers. In order to vary the experiment, half the flowers of both lots were on plants produced from self-fertilised seeds, and the other half on plants from crossed seeds. The sixty-four flowers with an excess of pollen yielded sixty-one capsules; and excluding four capsules, each of which contained only a single poor seed, the remainder contained on an average 5.07 seeds per capsule. The sixty-four flowers with only a little pollen placed on one side of the stigma yielded sixty-three capsules, and excluding one from the same cause as before, the remainder contained on an average 5.129 seeds. So that the flowers fertilised with little pollen yielded rather more capsules and seeds than did those fertilised with an excess; but the difference is too slight to be of any significance. On the other hand, the seeds produced by the flowers with an excess of pollen were a little heavier of the two; for 170 of them weighed 79.67 grains, whilst 170 seeds from the flowers with very little pollen weighed 79.20 grains. Both lots of seeds having been placed on damp sand presented no difference in their rate of germination. We may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.
One of the main points made in the last paragraph is __________.
Ipomoea purpurea produce smaller plants when under-pollinated.
The results of the experiment proved that the amount of pollen used causes significant variation in the number of seeds produced.
Mirabilis has a high rate of germination.
For heavier seeds, less pollen is required.
The experiment concerning the amount of pollen used to fertilize a flower proved that little or no error had initially been made in the main experiment.
The experiment concerning the amount of pollen used to fertilize a flower proved that little or no error had initially been made in the main experiment.
The author states that the purpose of the experiment discussed only in the fourth paragraph was initially to verify if the method of pollination used in the previous two seasons had caused or could cause an anomaly. At the end of the fourth paragraph, the author states that “we may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.”